Three New Species of the Genus Athemus (Coleoptera, Cantharidae) from Japan

13 Pages·2000·14.8 MB·English

by  Okushima, Y

Et、t''a, 「okyo,28(2):373-385,November15,2000
ThreeNewSpeciesoftheGenusAthemus
(Coleoptera,Cantharidae)fromJapan
YuichiOKUsHIMA
KurashikiMuseumofNaturalHistory,Chuo2-6-1,Kurashiki_shj
OkayamaProf.,710-0046Japan
and
Shoichi IMASAKA
EntomologicalLaboratory,FacultyofAgriculture,
KyushuUniversity,Fukuoka,812-8581Japan
Abstract ThreenewspeciesofthegenusAthe,mlsaredescribedfromJapanand
illuSt「ated. 0fthese,A.(Andlat/1e'nus),natsunaga1andA.(Andlathe,mis) tobirantlsare
CollectedfromtheJapanesemainland,andA.(At/1e,ne11us)naokil iscollectedfromthe
RyukyuIslands.
ThegenusAthemusLEWIS,1895,wasestablishedforaJapanesespecies,Te(lepho_
「uSSutu「e11uSMOTSCHULSKY,1860,andatthesametime,CantharisattristataKIEsEN-
WETTER,1874,wastransferredtolt.ManyJapanesespecieswerelatertransferredfrom
othergenera,oraddedasnewspecies,toAthemusbysubsequententomologists.Until
now, forty-one species ofAthemus havebeen recorded from Japan including the
RyukyuIslands(TAKAHAsHI,1998).However,sometaxonomicalproblemsstillremajn
aboutJapanesespecies.
Inrecentyears,wehavereexaminedtheJapanesespeciesofAthemuspreviously
recordedaswereunidentified,and foundthatat leastthreeofthemarenew toscience.
Ofthese,twowerecollectedfromtheJapanesemainlandandshouldbelongtothesub-
genusAndrathemusWITTMER,1978,whiletheotheronewascollected fromthesouth-
ern part of the Ryukyu Islands and should belong to the subgenus Atheme11us
WITTMER, 1972 (IMASAKA & YAMAJI,1989; IMASAKA & ABIRU, 1989; IMAsAKA &
NAKAMURA,1993;anonym,1997).Theywillbedescribedinthepresentpaper.
WewishtoexpressourheartythankstoDr.Shun-IchiUENooftheNationalSci-
enceMuseum(Nat.Hist),Tokyo, forhiscritical readingof theoriginalmanuscript,
andtoDr. MasatakaSAToofNagoyaWomen'sUniversity for hisconstantguidance
and co-operation to our studies. IMAsAKA also thanks Dr. Katsura MoRIMoTo of
Kyushu University for his continuous guidance. 0ur thanks are also due to Dr.
374 YuichiOKUsHIMAandShoichiIMASAKA
KyoichiroUEDA(KMNH), Messrs. MartinBRENDELL (BMNH), Malcolm KERLEY
(BMNH), KazuhiroTAKAHAsHl (KTC),MasamiMAsUMoTo(NIAES),Naoki TAKA-
HAsHl(NTC),andMs.HiromiNlwA(=H.URUsHIHARA,OMM) fortheirkindsupport
inloaningspecimensfromtheirinstitutionsorprivatecollections,andtoallthecollec-
tors, in particular Messrs. Yoshiaki MATsUNAGA, 0samu YAMAJI and Hirofumi
HAYAKAwA,whokindlyofferedinvaluablespecimenstous.
Thetypeseriesofthenewspeciestobedescribedinthispaperaredepositedin
thefollowinginstitutionsandpersonalcollections,whicharereferredtointhetextby
thefollowingabbreviations:BMNH:TheNaturalHistoryMuseum,London;KMNH:
Kitakyushu Museum and Institute of Natural History; KUF: Kyushu University,
Fukuoka;KURA:KurashikiMuseumofNaturalHistory;NIAES:NationalInstituteof
Agro-Environmenta1Sciences,Tsukuba;NWU:NagoyaWomens' University;OMM:
Omogo Mountain Museum; KTC: Kazuhiro TAKAHAsHI's collection; NTC: Naoki
TAKAHAsHl'scollection:SIC:Shoichi IMASAKA'scOllection.
Athemus(Andrathe″Ms)matsunagaiIMASAKAetOKUSHIMA,spnov.
[Japanesename:Matsunaga-jokai]
(Figs.1-4,7-12,Tablet)
Athe,mlssp.[Matsunaga-jokai]:IMAsAKA&ABIRU,1989,KumamotoKonchti-Dokokaiho,35(1):12,fig.
2-8. - IMAsAKA&NAKAMURA,1993,Misc.Rept.HiwaMusnat.Hist.,(31):51,pl 2, fig 27,pl.
4, fig 27. - IMAsAKA&0HTsUKA,1996,KumamotoKonchu-Dokokaiho,40(3);49, figs3-23,5-
23.
Male. Bodymostlyblackishbrownorblack;anteriorareabeforeeyes,mouth
parts,basalpartsofantennae,circumferenceofpronotum,prosternum,narrowventro-
lateralsideofeachelytron, forefemoraandtibiawithoutoutermargins,basalhalves
ofmiddleandhind femorawithout outermargins, andjointsof legs yellow; apical
partsofmandiblesandclawsreddishyellow.Eachelytrono量enwithayellowstripe
fromhumerustomiddlearea,developmentofwhichisvariablewithindividuals.Body
closelycoveredwithfinepalepubescence;apicalmarginofclypeusandlateralmar-
ginsofpronotum fringedwithpalebristles;eachelytronprovidedwithintermingled
palebristlesinadditiontoprimarypubescence,thoughtheybecomemuchsparser in
anteriorhalf.
Bodyveryslender.Headslightlyshorterthanitswidth;dorsumdepressedalong
theapicalmarginofclypeusandinlateralareasbeforeeyes, faintly depressedalong
themid-line;surfacesmoothwithfaintlustre,closelywithminuteandindistinctpunc-
tures;clypeusarcuateatapicalmargin;eyes large,globularandstrongly prominent,
Figs. 1-6. Athe'mlsspp fromJapan. - 1-4.A. (And'athe,1tls) ,11ats″l1agal' IMAsAKAet OKUsHIMA,
sp nov; 1, (holotype), from Hiroshima Prof.; 2, (allotype), from Hiroshima Pref; 3,
,
(paratype), fromOkayamaPref;4, (paratype),fromShizuokaPref. - 5.A.(A,td,at/1emus)to-
,
biranusOKUSHIMAet IMAsAKA, sp nov., (3(holotype), fromNagano Pref. - 6. A. (Athetne11us)
naokiiOKUsHIMAetIMAsAKA,sp nov., (holotype),fromIriomote-jimaIs.
ThreeNewAthemusfromJapan 375
376 YuichiOKUsHIMAandShoichiIMASAKA
ratioofthediameterofaneyetointerocularspace1:1.5;labialpalpuswithSemiCi「Cu-
farapicalsegment;maxillarypalpuswithsomewhatslenderandknife-shapedapical
segment;antennaefiliformandslender,attainingtoapicalthirdofelytra,1stSegment
clavate,2ndshortandalittleexpandedapicad,3rdto11thsubcylindrica1,eachof4th
to10thsegmentswithalongitudinalgrooveonthedorso-externa1side,relativefen9ths
ofantennalsegmentsasfollows:- 18:10:14:18:18:18:18:17:16:14:17.
pronotumsubquadrate, faintly expandedposteria 0.80times(in theholotype;
range0.75-0.83)aswideashead,1.00(1.00-1.15)timesaslongaswide;anterior
marginarcuate;posteriormarginweaklyarcuate; lateralmarginssinuate;anterioran-
glesrounded;posterioranglesobtuse;discconvex,particularlysointhepostero-1atera1
areas,stronglydepressedalongtheposteriormargin, antero-1atera1areashollowed;
medic-longitudinal furrowindistinctinanteriorarea;surfacesmoothwith faintlustre.
Scutellumtriangularwithbluntapex.
Elytraconjointly155(1.50-1.61)timesaswideaspronotum,3.71(3.39-3.81)
timesaslongaswide,thesidessubpara11e1thoughslightlyconvexatbasal fourth;dor-
sumcloselyandrugoselypunctate,thoughweaklyinbasalpart;costaehardlyrecog-
nizableoneachelytron.
Legsconsiderablyslender;eachfemurmostlystraight;eachtibiamostlystraight
though feeblyarcuateat thebase;eachouter clawof foreandmiddlelegsprovided
withathumb-shapedtoothatthebase,theotherclawssimple.
Malegenitalia:- Ventralprocessofeachparamerestraightandslightlyleaning
inwards,eachdorsalplateroundedattheapex,innermarginfaintlyconcave.Eachlat-
erophysisslightlycurvedtowardstheapexofdorsalplate,withitsapexsubtruncated
andpointedatthetip.Innersaclengthenedbehin aslongastegmen(Figs 7-11).
Lengthofbody:7.55mm(intheholotype; range6.40-7.80,measured fromthe
anteriormarginofclypeustotheapicesofelytra);breadthofbody:1.55(1.30-1.60)
mm(measuredatthewidestpartofconjointelytra).
Female. Bodysomewhatlongerandwider thaninthemale.Apicalmarginof
clypeussubtruncated.Eyesnotsolargeasinthemale, ratioofthediameterofaneye
to interocularspace l:2.0.Antennaealittleshorter thanin themaleand lackinga
grooveoneachsegment.Pronotum0.83-0.96timesaswideashead,0.92-1.05times
as longas wide.Elytraconjointly 148-1.64timesaswideaspronotum,3.10-3.67
timesaslongaswide.Eighthabdominalsternitedeeplyemarginateoneachsideof
terminalmargin, forminglargeroundedlateral lobesandawidemedianlobe,thelatter
ofwhichisnotchedatthemiddleandroundedoneachside(Fig. l2).
Lengthofbody:6.95-8.65mm;breadthofbody:1.50-2.10mm.
Typeofcolour、レ,ariat1on. TypeB:eachelytronentirelyblackexcept fornarrow
ventro-1atera1sideasshowninFigs.1-2;typeBY:eachelytronwithsmallyellowish
markonhumerusasinFig 3;typeY:eachelytronwithayellowstripefromhumerus
tomiddleareaasinFig 4.
Typeseries. Holotype: (B) (TypeNo 3111,KyushuUniv),Juppo-zan-rindo,
Yoshiwa-mura, Hiroshima Prof., Honshu, Japan, 19-V-1989, Y. MATsuNAGA leg.
ThreeNewAthem1lsfromJapan
377
l2
Figs. 7-12. Athem11s(Andnathemls)matsunagaiIMAsAKAetOKUsHIMA,spnov. - 7-9,Malegenitalia
(6,ventralview;7,lateralview;8,dorsalview);10-11,medianlobewithlaterophysesofmalegeni-
talia(10,lateralview;11,dorsalview);12,8thabdominalsterniteinfemale.(Scale:0.5mm)
(KUF).Allotype: (B),Nakatsuya,Yoshiwa-mura,HiroshimaProf.,Honshu,Japan,
24-V-1988,S. IMAsAKAleg.(KUF).Paratypes: [Honshu,Japan]KanagawaPref:1
(Y),Tennoji-one,Mt.Tanzawa-yama,2-VII-1984,K.TAKAHAsHl leg.(KTC,paratype
of . okzlyugawaranus).ShizuokaPref: 1 (Y),Mt.Kurakake-yama, Kannami-cho,
30-IV-1991, H.NAGAoKA leg. (KURA); 1(5 (Y), Mt. Amagi-san, Izu Pen., 3-VI-
378 YuichiOKUsHIMAandShoichiIMASAKA
prefecturaldjslrjbutionofcolourpatternsoftheelytrainthetypeseriesofAt1/emuS('i'ld'athe-
Table 1.
,nus)matsunagaiIMASAKAetOKUSHIMA,SP・nov・
Honshu
Colourtypes
ofelytra Kanagawa Shizuoka Mie Kyoto Hyogo Nara Shimane
Y Male l
Female 5
BY Male
Female
B Male
Female
Honshu Shikoku
Colourtypes
ofelytra
Okayama Hiroshima Tokushima Ehime Kochi
Y Mafo 5
maIe 4
BY Male 3
maIe 1
B Mate 4
maIe 8
Kyushu
Colourtypes
ofelytra Nagasaki Kumamoto Oita Miyazaki Kagoshima Yaku-shima
Y Male 1
Female I 1
BY Male l 9
Female l 5
B Male l8 2 4
Female 29 2 8
l990,H.KoJIMAleg. (KURA); l (Y),Mt.Amagi-san,IzuPen.,8-VI-1991,H.NA-
GAoKAleg.(KURA); l (Y),1 (Y),Mt.Amagi-san,IzuPen.,8-VI-1991,nocollec-
tor'sname(KURA); l(i (Y),Mt.Amagi-san,IzuPen.,9-VI-1991,H.NAGAoKA leg.
(KURA).MieProf.: 1 (Y),Hirakura,Misugi-mura, 14-V-1967, H. IcHIHAsHl leg.
(NWU);19(Y),Hirakura,Misugi-mura,4 V-l987,T.IMAMURAleg.(NWU);1 (Y),
Hirakura,Misugi-mura,3-V-1988,K.AKITAleg. (NWU);1 (Y),Hirakura,Misugi-
mura,4-V-1988,M.SAITo leg. (SIC); l (Y),Hirakura,Misugi-mura,27-V-l988,
N. KANIE leg. (NWU); 1 (Y), Chichigatani, Miyagawa-mura, 5-V-l989, N.
NARUKAwA leg. (NWU). KyotoPref: 1 (Y), Ashiu,Miyama-cho, 6-VI-1991, K.
ThreeNewAt1/e,mlsfromJapan 379
MAsAKl leg. (SIC). HyogoPref: 3 (2Y, IBY), Akasai-keikoku, Haga-cho, 13-
V-1993,H.0KADAleg.(SIC);1 (BY),19(BY),Akasai-keikoku,Haga-cho,16-V-
I993,H. 0KADA leg. (SIC). NaraPref: 2 (Y),Mt. 0daigahara,Kamikitayama-
mura,29-VI-1975, K.MlzuNo leg. (SIC); 1 (Y),Mt.0daigahara,Kamikitayama-
mura,19-VI-1977,K.MlzuNoleg. (SIC);1 (Y),1e(Y),Mt.0daigahara,Kamiki-
tayama-mura,16-VI-1985, K.URATA leg. (SIC);1 (Y),Mt. 0daigahara,Kamiki-
tayama-mura,14-V-1986,N.NARUKAwAleg.(SIC);19(Y),Mt.0daigahara,Kamiki-
tayama-mura, 21-VI-1987, N. NARUKAwA leg. (NWU); I (Y), Mt. 0daigahara,
Kamikitayama-mura,26-VI-1992,K.MATSUMOTOleg. (KURA);1 (Y),Shirakawa-
mata,Kamikitayama-mura,4-V-1980,K. MIzuNo leg. (SIC);2 (Y),Mt. Misen,
OmineMts.,21-VI-1987,K.M1zuNoleg.(SIC);19(Y),Mt. 0bako-dake,Yoshino-
gun, 11-V-1986, T. HATAYAMA leg. (SIC); 4 (Y), Mt. 0bako-dake, Yoshino-gun,
1-VI-1986,T.HATAYAMAleg.(SIC).ShimaneProf.:19(B),Ai,Nita-oho,11-V-1960,
T.FUJIMuRAleg.(NIAES).OkayamaProf.:1 (BY), 0kutsugawa,Shohoku-cho,19-
V-1996,A. WATANABE leg. (KURA); 1 (BY),Shimotsugawa, Kame-oho, 11-V-
I997,0.YAMAJ1leg. (KURA).HiroshimaPref:1 (B), 1 (B),samedataasfor the
holotype(SIC);19(B),Nakatsuya,Yoshiwa-mura,27-V-1984,H.0KADAleg.(SIC);
4 (B),19(B),Nakatsuya,Yoshiwa-mura,23-V-1988,S.IMAsAKAleg.(SIC);8
(B),4 (B),samedataasfortheallotype(SIC).[Shikoku,Japan]TokushimaPref:
3 (Y), Mt. Takagi-san,Kisawa-son,15-VI-1986,A.WATANABEleg. (SIC); 2
(Y),1 (Y),Dosu-toge,Kisawa-son,17-V-1987, A.WATANABEleg. (SIC); 1 (Y),
Minokoshi,Higashiiyayama-son,20-V-1993,Y. 0KusHIMAleg. (KURA); l (BY),
Kuwadaira-Tsurugi-san,Ichiu-son,20-VI-1993,S.MANoleg.(KTC);1 (BY),Mt.
Tsurugi-san, Ichiu-son,20-VI-1993,S.MANoleg.(KTC);4 (2Y,2BY), Mt. Tsu-
rugi_san,alt. 1,600-1,900m,22-V-1998,M.YosHIDAleg. (SIC).EhimePref:2
(BY),Nibukawa,Tamagawa-cho,3-V-l996,M.SHIRAIsHI leg.(SIC);2 (B),Mt.
Narahara-yama,Tamagawa-cho,26-IV-l998,M.SHIRAISHI leg. (SIC); I (Y), 3
(Y),Joju-sha,Mt.Ishizuchi-san,26-VI-1994,M.SHIRAISHI le9. (SIC);1 (Y),Mt.
Ishjzuchi-san,27-V-1990,H.KusuNoKlleg.(OMM);19(Y),0damiyama, 0da-oho,
6_V-1995,N. 0HBAYAsH1leg. (NWU); l (Y), 0damiyama,0da-oho,6-V-1995,K.
AITAleg.(NwU).KochiPref:1(5'(B), forkofNaka-gawa&Nishi-kawa-Senbon-
yama_tozan-guchi, Umaji-mura, 2-V-1988, K. HAGA leg. (KTC); l (Y), Ten9u-
kogen,Hjgashjtsuno-mura,13-VI-1995,Y.0KUsHIMAleg.(KURA).[Kyushu,Japan]
Nagasakipref:4 (B),Mt.Unzen-dake,0bama-cho,12-VI-1986,S.IMASAKAle9.
(sIc);1 (B),2 (B),Mt.Unzen-dake, 0bama-cho,28-V-l987,S. IMASAKA Ie9.
(sic);3 (1BY,2B),3 (1Y,1BY,1B),Mt.Unzen-dake,0bama-cho,31-V-1989,
s.IMAsAKAleg.(SIC).KumamotoPref:5 (1BY,4B),7 (B),Shiiya-tOge,Yabe-
cho, 6_vI_1g8g, S. IMAsAKA leg. (KURA); 2 (B), Shiiya-tOge, Yabe-Cho,
6_vl_1gg5,s.IMAsAKAleg.(SIC);19(B),Naidaijin-kyo,Yabe-cho,12-V-1984,S・
(B),Hagi,Izumi-mura,19-V-1996,S.IMASAKAle9・(KURA);
OGATAleg. (SIC); 1
3 (B),2 (B),Hagi,Izumi-mura,4-V-1997,S.IMASAKAleg.(SIC);2 (B),Mt・
Hakucho_zan, Izumi-mura,17-VI-1984,S. IMASAKAleg. (KURA);4 (1BY, 3B),
380 YuichiOKUsHIMAand ShoichiIMAsAKA
Mt. Hakucho-zan, Izumi-mura, 18-VI-1984,S. IMAsAKA leg. (SIC);4 (B),9
(8B,1Y),Mt.Hakucho-zan,Izumi-mura,5-V-1998,R.NoDAleg.(SIC);4 (B),1
(B),Mt.Shiraga-dake,Ue-mura,31-V-1994,S. IMAsAKA leg. (SIC);2 (B),3
(B), Mt. Shiraga-dake, Ue-mura, 17-V-1997,S. IMAsAKAleg.(SIC).0itaPref: I
(BY),Taisen-rindo,Mt.Hiiji-dake,18-V-1982,S.SAsAKl leg.(SIC); 1e(BY),Mt.
Kuro-dake,Shonai-cho,10-V-1983,Y.TAKAKURAleg.(KMNH);1 (BY),Mt.Kuro-
dake,KujuMts.,9-V-1987,S.0GATAleg.(SIC);1,3(BY),4 (3BY, IB),Mt.Kuro-
dake,KujuMts.,16-V-1989,S. IMAsAKAleg.(SIC);2 (1BY, IB),6 (4BY,2B),
Mt.Kuro-dake,KujuMts.,20-V-1992,M.NlsHIDAleg. (SIC);2 , (1BY, IB),Mt.
Kuro-dake,KujuMts.,25-V-1996.S. IMAsAKAleg.(SIC); 5 (1Y,1BY,3B),1
(Y),Taisen-rindo,KujuMts.,7-V-1989,M.NIsHIDAleg.(SIC);2 (B),8 (5BY,
3B),Mt.Katamuki-yama,30-IV-1990,S.SAsAKI leg.(KURA);4 (3BY, IB), l
(B), Hoei-rindo, Mt. Katamuki-yama, 26-IV-1998,R.NoDA leg. (KURA); 6
(1BY, 5B), 4 (B),Hoei-rindo, 0gata-machi,29-IV-1999,R.NoDAleg.(SIC);1
(BY),Mt.Soho-san,24-V-1995,N.TAKAHAsHIleg.(NTC).MiyazakiPref:1 (BY),
Mt.GOyo-dake,Hinokage-cho,15-VI-1996,S. IMAsAKAleg.(SIC);1 (B),Mt Shj_
raiwa-yama,Gokase-cho, 18-V-1991, A. NAGA1leg. (SIC); I (B), Ebjno_kogen,
Ebino-shi,24-V-1990,A.NAGAlleg.(SIC).KagoshimaProf.:5 (B),2 (B),Mt
Shibi-San, Miyanojo-cho, 14-V-1991,S. IMAsAKAleg. (KURA,SIC); 1 (B), Mt
Ki「iShima-yama, 13-V-1991,S. IMAsAKAleg.(SIC);2(S(; (B),Yodogawa_rjndo,alt
1,300m, Yaku-ShimaIs., l7-V-1988,Y.MATsUNAGAleg.(SIC); 2 (B), 6 (B).
YOdOgaWa-gOya, Yaku-shima Is., 7-VI-1988, S. IMAsAKA leg. (SIC); I (B), Yo_
do9aWa-goya,Yaku-shimaIs.,9-VI-1988,S.IMAsAKAleg.(SIC);1
(B),Shjratanjun_
Sui-kyo-Tsuji-no-toge,Yaku-shimaIs.,4-V-1998,N.TAKAHAsH1leg.(NTC) [Nodo_
taileddatal1 (Y),“Japan./G.Lewis./1910-320.”(BMNH).
Distribution. Japan:Honshu(westofKanagawaPref),Shikoku,Kyushu,Yaku_
shima Is.
Notes. ThisnewspeciescloselyresemblesA. (And,at11emus)okMyugawa,・anus
TAKAHASHI,1992fromKanagawaProf.,Honshu,particularlyintheyellowstripedtype
(Y),butCanbedistinguished fromthelatterbythecolourofthemiddleandhjnd
feme「aWhichareblackishalongtheoutermargins,andthestructureofthemalegenj_
talia,aboveall inthebroaderdorsalplates.Theblackishtype(B)canbeeasilydist1n_
9uiShed f「om A. (Andi'athemus) okuyugawalamtsTAKAHAsHl by the characterjstjc
colour
TheColOu「ofthebodyshowsatendencytobecomeblackishinthesouthwestern
a「eaofthedistributionalrangeofthespecies,andtobecomeyellowishintheeastern
a「ea(Tablet).However,thismerelyshowsgeographical variationwjthjnthesame
Species,becauseitiscontinuousandnomorphologicaldifferencehasbeenobserved
except forcoloration.
Ithasbecomeclearbyourreexaminationthatoneoftheparatypesof
(/n_
Hgeftlj duatﾝenZlISoた) t:yugawa' 'ams TAKAHAsI actualy belons o ths nw specje
ThisSpeciesSi Collectedonvariousftowers(Palulac11inensisKolDz
var pj1osa
ThreeNewAt1/e,mlsfromJapan 381
NAKA1,CornuscontroversaHEMsLEY,etc.) intheupper lucidophy11ousto thelower
cool temperatedeciduousbroadleaved forests.Livingindividuals1ooklikeEncyc1ops
olivaceaBATES,a1ongicornbeetle,on forestflowers.
ThespecificnameisgiveninhonourofMr.YoshiakiMATsuNAGAwhocollecteda
partofthetypeseriesincludingtheholotypeofthisinterestingnewspecies.
Athemus(Andmthemus)tobiramlsOKUsHIMAetIMAsAKA,sp nov.
[Japanesename:Tobira-jokai]
(Figs5,13-15)
Athemtlssp. [Tobira-jokai]:anon.,1997, Matsumoto-shino Konchu, Matsumoto-shishi Shizen-bumon
Chosa-hokoku-sho,Matsumoto-shi,1,p 47.
Male. Bodymostly yellow.Eyes black;mandiblesand claws faintly reddish;
antennae, tarsi,metasternumandabdominalsternitessomewhat dusky. Body closely
coveredwithfineyellowishpubescence;apicalmarginofclypeusfringedwithyellow-
ishbristles;eachelytronwithintermingledyellowishbristlesinadditiontoprimary
pubescence.
Bodyveryslender.Headslightlyshorterthanitswidth;dorsumdepressedalong
theapicalmarginofclypeusandinlateralareasbeforeeyes;surfacesmoothwithfaint
lustre,sparselywithminuteandindistinctpunctures;apicalmarginofclypeusarcuate
withitscentrefaintly indented;eyeslarge,globularandstronglyprominent,ratioof
thediameterofaneyetointerocularspace1:1.5;labialpalpuswithsemicircularapi-
calsegment;maxillarypalpuswithsomewhatslenderandknife-shapedapicalseg-
ment; antennaefiliformandslender,attainingtoapical thirdofelytra, 1stsegment
clavate,2ndshortandalittleexpandedapicad,3rdto11thsubcylindrica1,eachof4th
to10thsegmentswithalongitudinalgrooveonthedorso-externa1side,butthatonthe
4thisshort,relativelengthsofantennalsegmentsasfollows:- 15:10:12:14:17:16:
16:16:16:14:15.
Pronotumsubquadrate, faintly expandedposteria 0.87 times as wide as hea
0.g6tjmesaslongaswide;anteriormarginarcuate;posteriormarginweaklyarcuate;
lateralmarginssinuate,weaklyhollowedbehindanterioranglesandconstrictedjust
beforeposterior angles; anterior anglesrounded; posterior angles rectangular and
slightlyprojected;discconvex,particularlysointhepostero-1atera1areas,stron9lyde-
pressedalongtheposteriormargin,antero-1ateralareashollowed;medic-longitudinal
furrowdistinctonlyincentralarea;surfacesmoothwithfaintlustre.Scutellumt「Ian-
gular,withbluntapex.
Elytraconjointly154timesaswideaspronotum,3.4timesaston9aswide,the
sjdessubpara1le1thoughslightlyconvexatbasal fourth;dorsumcloselyandrugosely
punctate,thoughweaklyinbasalpart;eachelytronprovidedwithtwovagueCostae.
Legsconsjderablyslender;eachfemurmostlystraight;eachtibiamostlystraight
though feeblyarcuateatthebase;eachouterclawof foreandmiddlele9SP「oVided
withathumb-shapedtoothatthebase,theotherclawssimple.
382 YuichiOKUsHIMAandShoichilMAsAKA
Figs. 13-15. MalegenitaliaofAthemus(And'athe1mts)tobiran11sOKusHIMAetIMAsAKA,sp nov;13
ventralview;l4,lateralview;15,dorsalview.(Scale:0.5mm)
Malegenitalia:- Ventralprocessofeachparamerestraightandleaninginwards,
eachdorsalplateroundedattheapex,slightlyexpandedinwards.Eachlaterophysis
curvedtowardstheapexofeachdorsalplate,pointedatthetip. Innersaclengthened
behin aslongastegmen(Figs.13-15).
Lengthofbody: 9.0mm(measured fromtheanteriormarginofclypeustothe
apicesofelytra);breadthofbody:2.0mm(measuredatthewidestpartofconjointely_
tra).
Female. Unknown.
Type specimen. Holotype: (Type No 3112, Kyushu Univ), Tobjra, Ma_
tSumOto-Shi,NaganoPref.,Honshu,Japan,12-VI-l995,H.HAYAKAwAleg.(KUF).
Distribution. Japan:centralHonshu(NaganoPref).
Notes. This new species somewhat resembles A. (Andrathemus) okuyu_
9awaranusTAKAHASHI,1992fromKanagawaPref.,Honshu,butcaneasilybedistin_
guished fromthelatterbythewhollyyellowbodyandthedorsalplatesofthemale
genitaliaslightlyexpandedinwards.