Table Of Content九州大学学術情報リポジトリ
Kyushu University Institutional Repository
Study on the Tribe Ochyromerini (Coleoptera,
Curculionidae) from East Asia I, Descriptions
of New Species of the Genera Endaeus and
Endaenidius
Kojima, Hiroaki
Entomological Laboratory. Faculty of Agriculture, Kyushu University
Morimoto, Katsura
Entomological Laboratory. Faculty of Agriculture, Kyushu University
http://hdl.handle.net/2324/2582
出版情報：ESAKIA. 35, pp.63-102, 1995-01-31. 九州大学農学部昆虫学教室
バージョン：
権利関係：
,AIKASE :)53( ,201-36 yraunaJ 31, 5991 63
Study on the Tribe Ochyromerini (Coleoptera, Curculio-
nidae) from East Asia I, Descriptions of New Species
of the Genera Endaeus and Endaenidiusl)
ikaoriH KOJIMA dna arustaKM ORIMOTO
lacigolom o,tynrEotarobaL ytlucaF fo,erutlucirgA
,uyh ts,iuasykrKoeuvkiunFU 8 21napaJ
Abstract sihTsi the first part of our systematic study on the tribe Ochyromerini
)inieadnE=( morf tsaE aisA htiw snoitpircsed fo 82 wen seiceps fo eht sliveew nieht
genera Endaeus Schoenherr and Endaenidius Morimoto as follows: Endaeusalbo .psl .voni .E( ,)aisyalnaM eatus Endaeus niger .ps .von .E( ,)aisyalaM Endaeus
zonatus .ps nov. (E. Malaysia), Endaeus robustus sp. nov. (Thailand), Endaeus
longipes .ps .von .E( ,)aisyalaM Endaeus formosanus sp. nov. (Taiwan), Endaeus
elongatus .ps nov. (Taiwan), Endaeus sajapi sp. nov. (W. Malaysia), Endaeus
selangorensis sp. nov. (W. Malaysia), Endaeus makiharai sp. nov. (Indonesia),
Endaeus momosei .ps .von .E( ,)aisyalaM Endaeus giganteus .ps .von .E(,)aisyalaM
sp. nov. (E. Malaysia), Endaeus malayanus sp. nov. (W.
Malaysia), Endaeus bicoloripennis .ps .von .E( ,)aisyalaM Endaeus nigripes .ps .von
.E( ,)aisyalaM Endaenidius babai .ps .von ,)nawiaT( Endaenidius okushimai .ps.von
(W. Malaysia), Endaenidius enicosanthi sp. nov., (E. Malaysia), Endaenidius
longirostris .ps .von ,)dnaliahT( Endaenidius macrommatus .ps .von .E(,)aisyalaM
Endaenidius maetoi .ps .von .W( ,)aisyalaM Endaenidius sarawakensis .ps .von.E(
Malaysia), Endaenidius kemae .ps .von .E( ,)aisyalaM Endaenidius crassipes .ps.von
.E( ,)aisyalaM Endaenidius spinipes .ps .von .E( ,)aisyalaM Endaenidius polyalthiae
.ps .von .E( )aisyalaM dna Endaenidius brevirostris .ps .von .E( .)aisyalaM syeKot
seiceps detaert ni siht ,repap eht enil snoitartsulli fo sutibah dna tnatropmi serutaefera
provided.
The tribe Ochyromerini contains small-sized weevils and predominantly occurs in the
laciport saera fo eht latneirO dna naipoihtE .snoiger yehT era yllausu derutpac no ehtsrewolf
dna sevael yb gnitaeb dna ,gnipeews dna emos era netfo detcartta ot ,thgil tub tsom fo meht era
erar dna ylno a wef slaudividni evah neeb detcelloc.yliratnemgarf
traP fo eht sliveew detaert ni siht yduts erew derutpac gnirud eht syevrus fo ehtytisrevidoib
ni eht laciport niar stserof detcudnoc yb eht smaet fo eht otoyK ytisrevinU dna eht yrtseroFdna
tseroF stcudorP hcraeseR.etutitsnI tsomlA lla fo eht sliveew derutpac ni eht seiponac era foeht
)1 noitubirtnoC morf eht lacigolomotnE ,yrotarobaL ytlucaF fo ,erutlucirgA uhsuyK,ytisrevinU
akoukuF .reS( .oN ,4 .oN.)97
64 H. KOJIMA & K. MORIMOTO
debircse dsneuiceps dna nwonk seiceps evah neeb derutpac ylno no eht srewolf dna sevaelfo
sr s. t ege ss sewnedti tehoir n nhscgtlvtfata Teag ioalyif uslepncnseraei i gmpepfshootneahct
.sc is ptneosihr ettnrioafr
Present study has been carried out for the purpose of clarifying the phylogenetic
spihsnoitaler fo eht detaler taxa, dna snoitpircsed fo wen seiceps fo eht areneg and
Endaeus
erofeb su era detneserp ereh sa eht tsrif trap fo ruo .ydu ttsnemtaerT foesenapaJ
Endaenidius
seiceps si won gnitnirp erehwesle ni eerht.strap
erofeB gnio g,rehtruf ew hsiw ot sserpxe ruo ytraeh sknaht ot eht etal .rD .K ,abaB .rM.0
.K .rD ,otomiK .S .forP ,aturuF ,oteaM .rM ,otomustaM .K .rM ,arahikaM .H .rM ,egakiM .T .rM
.S ,awakayiM .rD .Y ,ekatayiM .rM .K ,esomoM .rD .T ,enakaN .rM .Y ,amihsukO .forP.T
..f T odirnjPa a .s.fa HoS,raPsugiaS uzorihS .slai r gesntsi aer nm nerirdfionfefiohkt
Descriptions
suneG Endaeus Schoenherr
: seepiycTeps rrehneohcS
Endaeus castus
am iejeoSK & .sis os nedgcnanaied r,eyfme y r) no srotnsonoief ym(rsiproM
Key to species treated in this paper
)8(l d eeytt l ane rptliatornuoiaao gpbhreni sFhatxtep a.freoelngnnia
)5(2 Body oblong ovate, blackish. Antenna1 seborts gninnur ot htaeneb eht esabfo
.murtsor
ydoB tahwe m.orsen nxiahrtohtor Ptuoba 4.1 semit sa ediw sa ,gnol ylkaewdednuor
3(4)
.yllaretal artylE 44.1 semit sa gnol sa .ediw . . . . . . . . . 2. .ps .von
niger
art y.lyEllare tdaeldn u,ogr neso dalssieaw m 3ix.ta1roht o.rrPetsu btoarhwe myodsoB
4(3)
63.1 semit sa gnol sa .ediw . . . . . . . . . . . . . . . . . . . . . .1 .ps .von
albolineatus
W) yd oyBld a,hoesrtibaw vonolwloeryb ot kra d. n1waonrnbetnA seborts gn iontneuhrt
.seye fo trap rewol
murtsoR a elttil retrohs naht .mutonorp mreD hsiwolley ,nworb laretal snigramfo
h(7)
,mu tloanroe rmspuehlgna fo artyle d nm au.lhlseitkuccaslb . . .4. .ps.von
robustus
murtsoR hcum regnol naht .mutonorp mreD krad ,nworb htiw eannetna dnasgel
7(c)
hsiwolley .nworb artylE htiw owt esrevsnart sdnab fo hsiyerg etihwtnebmucer
mrofites selacs neewteb nedlog hsinworb tnebmucer esotes.saera
. . . . . . . . . . . . . . . . . . 3. ..vposn
zonatus
)l(8 d e edt anntiuoab orierF thxtep afr.oeengndie
xarohtorP ht iswuo uncoiipsssneor cpdeedllif htiw eates dniheb eht roiretn anigramni
9w
1an n.eetlnaAmef seborts .mur t es f soeoharhtbta e gnonetibn nur
)11(Ol eh a tti.inbewoaitrtaFt nae r dto. in mrsboheeurcfhoFatueog i.steentyoaE cgyndoolBe
.esab eht raen nigram renni eht ta eates tcere wef
. . . . . . . . . . . . . . . . . 6. .ps.von
elongatus
NEW ASIAN ENDAEUS AND ENDAENIDIUS 65
)01(ll seyE ton .detamixor pxpaarohtorP yltnedive tnereffid ni htob ,sexes tsediw talacipa
).e el dtarramoite h nf atd e(.er- nad) loeeiieetmnlhnlhteodeuatiFdbmli(m
. . . . . . . . . . . . . . . 5. .ps.von
formosanus
12(9) ndoni i.srt xhnsoueaeieiohrbgrrhtorpethaetitmdnwoarP
13(22) e hattinib owiar tFwo rfo tcerebus e arteetsu ontasht eht tce reeates fo aromef tarenni
.egde
14(21) .x e eprha atyelnlanre tdneita leiadi btintorF
15(20) .suo r1oalnonceitBnA seborts gn i o nrttenerhu w.atrfospoleye
16(19) .nworb hsidder ro hsiwolley mulletucS
17(18) artylE dedis-lellarap ta lasab .flah eannetnA ,hsikcalb gnitpecxe epacs dna tsrif
tnemges fo .elcinuf . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8. .ps .von
sajapi
18(17) . nhws oi terdth abdsgan eeirn.trdltee aiytlewlndhEAdtim . .9. .ps.von
selangorensis
19(16) mulletucS .hsikcalb . . . . . . . . . . . . . . . . . . . . . . . . . 10. .ps .von
makiharai
)51(02 ,suorolocnoC thgil hsidder .nworb 1annetnA seborts gninnur ot htaeneb eht esabfo
.murtsor . . . . . . . . . . . . . . . . . . . . . . . . . 11. .ps .von
momosei
)41(12 tnorF eaibit detalid yllanretni a elttil erofeb eht .elddim . . . . . 12. .ps.von
giganteus
)31(22 tnorF eaibit tuohtiw retuots.eates
)03(32 rsgoan osmlaurts o.Rhtf isgfan ostlanemg ehstx ihst iewlcin u1fannet n.AsuorolociB
.muton on rarphettr oehlst tail
)52(42 .xe pe ahtytallanret ndietal iydlka eewaib ittno r.Fhsikca lyblerit nseg ed lnaartylE
. . . . . . . . . . . . . . . . . 16. nigripes ..vposn
25(24) .nworb hsidder ot nworb hsiwolley sgeL
)92(62 .strap lacipa nworb hsiwolley rof tpecxe ,hsikcalb artylE
)82(72 seyE ylgnort sxevnoc morf .selp m1eatnnetnA seborts gninnur ot htaeneb eht esabfo
.murtsor artylE l-54.1 .50 semit sa gnol sa .ediw . . . . . 13. .ps.von
flavoapicalis
)72(82 sey Exevno cm o. rs1fealnpnmeettnA seborts gnin noutr e hrte wtorlap f o.asretyyelE
1.30-l .35 semit sa gnol sa .ediw . . . . . . . . . . . . . . . 15. .ps .von
bicoloripennis
29(26) artylE yleritne .hsikcalb . . . . . . . . . . . . . . . . . . . . . . . 14. .ps .von
malayanus
)32(03 ,suorolo chnsoiCdder .nworb 1annetnA elcinuf htiw htxis tnemges retrohs naht.htfif
murtsoR a elttil regnol naht .mutonorp . . . . . . . . . . . . . . 7. .ps .von
longipes
1. Endaeus albolineatus .ps.von
.giF()1
.elaM mreD ,kcalb lacipa trap fo ,murtsor 1annetna epacs dna tarsi .hsinworb
daehero Fneewteb seye ,worran 8/l semit sa ediw sa eht esab ,fmou rhttsiowert innaifedni
na i.daeemv osfe yxEevn o mc.o srmeful rp,tmysenotiRhs a el trteigln o,nlmauhtt ohntoirwpa
anirac no hcae edis gnola 1annetna ;eborts 1annetna seborts gninnur ot htaeneb eht esabfo
ylr a,esn egyneihc aeepr a;cmsurt sefolord d eiehmrto feelb tt t saidulejtre senainne t.nmAurtsor
sa gnol sa ;elcinuf elcinuf htiw tsrif tnemges tsuj a elttil regnol naht ,dnoces driht othtxis
lauqebus ni ,htgnel 3/2 semit sa gnol sa ;dnoces bulc ,citpillebus 9.2 semit sa gnol sa,daorb
.de n y ielsb lrtms gcanoa fn ie ecloornnsmaluuagsofeafsb
eht ta detcirtsnoc ,yllaretal dednuor ,elddim eht ta tsediw ,gnol sa ediw sa semit 3.1 xarohtorP
66
H. KOJIMA & K. MORIMOTO
Fig. 1. Endaeus albolineatus sp. .von - ,A ;elaM ,B dae h; xdanraoh t,ocrptnorf
;gel ,D.annetna
;xepa musrod ylesogur ,derutcnup htiw hsiyerg tnebmucer .eates nulletucS,ralugnairtbus
yle sdneehdtol chti whsiye regt imhrwo f.isteel saacrstylE 63.1 semit sa gnol sa ,edi wtsediwa
eltti ldniheb eht ;elddi mtsr ilfavr eytlneis ndeedhtolc htiw hsiyer geti hmwrofi tseeslacs sano
tnebmu che trdiewht ohl ccsaleavre trnei h,taort yenlpoei rltasnidutig ngonlim r do,nfmaulletucs
hsiy eeragtes y ldrnaaluge rari elt triel ghnsoil y.eeragte ssg edLehto l hcttniewhbsmiuyceerrg
a h;r gdthornn itma oiawo elh lhrotfut;otngie m onwapearatrffiearshts tcerebus egantoeelhsat
roir e;tnsiogpra mtnorf eaibi tylne v,edev rd ueyctlallaindretn ita eht xepa dna detniop tarenni
e keillk cdiespr aghh ntsaeoialwi bdi nt;ix h eefploagna sucnu . xteeahpta
Prosternum with coxae located in the middle, the distance between the coxae and
lani gsruacmlbuuss sa taerg sa tah tneewte bsiht suclus dn aroi r.e nteindgairsadrmeehdtnoUlc
.eates hsiyerg tnebmucer htiw
.nwonk n.UelameF
:htgneL 8.2.mm
:epytoloH elam epyT( .oN ,4692 uhsuyK ,).vinU ulabaniK .P .H ,.Q ,habaS oenroB ,.sI.E
.gel amijoK .H ,3991 .iii .81-61 ,aisyalaM
.E aisyalaM.)habaS(
Distribution.
sihT seiceps si ralimis ot ssoV te 6jjuhC morf napaJ dna
Remarks. E. nipponicus E. ater
llahsraM morf ,aidnI tub elbarapes morf meht yb eht gniwollof :stniop tnorf eaibitylprahs
detnio pta eht xepa fo re n,neilg ndan iahrom ehftiw gno lea t regos ninorilnegaetresawomtpeb
ekil-elk cpir sahg htneasioawli bd i ndthinth a o edoeh ntsteaahbt sucnu .el ae mhnti
information.
Biology. No
WNEANISA ENDAEUS DNA ENDAENIDIUS 67
2. Endaeus niger .ps.von
)2 .giF(
dna eaibit fo trap lacipa ,kcalb mreD .elameF tarsi .hsin worb
dnaeee hwsetereyobeF l/4 se m sieatd isw. as ee x .ehlmesmstpoveu amrnyrfbefoEtotcsor
mu r,ty snroieRhgsn onmlauhttono r4p( : 3), g nydilrwkaoawre orwetahnt xepa nliar,ewteailv
tuohtiw naidem ;anirac 1annetna seborts gninnur ot htaeneb eht esab fo .murtsoreannetnA
detresni ta eht elddim fo ;murtsor epacs gnihcaer ,seye a elttil retrohs naht elcinuf 3( : 3.5);
elcin uhftiw ts rtinfem gteusoba 5.1 semit sa gnol sa ,dno cdersiht 6.0 semit sa gnol sa,dnoces
hh ttorxtuiosf lauqebus h c,ahetg nneil 413 semi tsa gno l , ec sci;aitdrp r hibistluhalltecgbnuosl
.denib meolcci nsfutofnem geees rlha t sg s an,asbodala osrab
xarohtorP 5/4 ylka e,wyllaret adlednu oyrlka e,wxe pe ahtdtaetcirtsn o,cgn oselad isswaem it
detanigrame ta eht elddim fo eht roiretna;nigram musrod ylgnorts derutcnupgnitpecxe
,.h reshaaitltyiueewgsrnga imrutl bl.h ueest st aind urtye e,coeebhhtSisrmttrrguioaecwlptecnra
artylE 44.1 semit sa gnol sa ,ediw tsediw ta eht ;elddim hcae lavretni htiw hsiyertgnebmucer
.eates sgeL dehtolc htiw tnebmucer hsiyerg ;eates tnorf aromef htiw a prahs lausuralugnairt
;htoot tnorf eaibit devruc ,ylneve detalid yllanretni ta eht xepa dna detniop ta renni elgnafo
.xepa
mumetsorP htiw eaxoc detacol ni eht ,elddim eht ecnatsid neewteb eht eaxoc dna-bus
h tdiewht oeldcisre d.nnUig rraomire tdsnnuaac ls uinshetew tt tea sab hsaesturac glluasnigram
.h estaintyeeebsrmgucer
Fig. 2. Endaeus niger .ps ,.von - ,;Aelam e,FB da e;hx adrnoa h,tCo rtpnor;fgel
,D.annetna
68 AM I.JHOK & O T.OKMIROM
.nwonknU .elaM
:htgneL 58.2.mm
:ep yetloalmo eHefpyT( .oN ,56 9u2h s,u)y.KvinU 61 seli mtniop m ourafsginnianmeiKK(
.gel amijoK .H ,3991 .iii .7 ,aisyalaM .E ,oenroB ,habaS ,)daor
.)ha baaiSs (y. an.loEaiMtubirtsiD
Remarks. si hsTeiceps si yrev esolc ot E. albolineatus .ps ,.von tub exhatroht odrnpaartyle
.renniht elttil a era
.ygol.oni oBoiNtamrofni
3. Endaeus zonatus .ps.von
.giF()3
elaM dna .elamef mreD krad ,nworb netfo yltrap ,hsikcalb ,murtsor eannetna dnasgel
hsiwolley.nworb
nsee eydweatee hb. e ehryasoetiFrewg sotlnleeb y mdhuet chidetwaroelHc 312 e dssieaw mit
sa eht esab fo ,murtsor dehtolc htiw tnebmucerbus hsiwolley yerg ,eates htiw naetinifedni
naid e.maevof seyE xevnoc mo r.fse lmpumret tsroeRgnol na hmtutonor p01( : 7 ni elam ro11
: 7 ni ,)elamef htiw a naidem dna owt swor fo tniaf eanirac no hcae edis dniheb eht1annetna
;noitresni 1annetna seborts gninnur ot eht rewol trap fo .se yeean ndeettnrAesni ta lacipa -eno
htruof )elam( ro a elttil dniheb lac idpraih t)-eelnaomef( fo ;murtsor epacs g nt,ioshnecyaeera
el trteigln on laehltcinu f3( : ;) 4elcin uhftiw ts rtinfemges sa gnol sa ,dno cdersiht 6.0 semitsa
Fig. 3. Endaeus zonatus ..vposn - ,;Aelam e,FB da e;hx adrnoa h,tCo rtpnor;fgel
,D.annetna
NAI SWAEN ENDAEUS DNA ENDAENIDIUS 69
gnol sa ,dnoces htruof ot htxis lauqebus ni ,htgnel 7.0 semit sa gnol sa ;driht bulc 68.2 semitsa
gnol sa ,daorb lasab owt stnemges fo elcinuf.denibmoc
Prothorax 1.4-1.5 semit sa ediw sa ,gnol tsediw a elttil dniheb eht ,elddim dednuor,yllaretal
ylkaew detcirtsnoc ta eht ;xepa musrod ylenif ,derutcnup dehtolc htiw hsiwolley yergtnebmucer
setiform scales and medially greyish white setiform scales and faintly forming a longitudinal
stripe. Scutellum tongue-shaped, clothed with recumbent greyish white setiform scales. Elytra
yldaorb ,etavo 2.1 semit sa gnol sa ,ediw tsediw a elttil dniheb eht ,elddim htiw noitalucamsa
:swollof tnebmucer hsiyerg etihw mrofites selacs gnimrof owt esrevsnart sdnab raen eht esabdna
a little behind the middle, remaining area clothed with golden brown recumbent setae; each
lavretni htiw a wor fo tnebmucerbus nedlog nworb regnol dna retuots .eates sgeL dehtolchtiw
tnebmucer hsitihw ;eates tnorf aromef htiw a lausu ralugnairt ;htoot tnorf eaibit ylneve,devruc
dilated internally at the apex, and weakly angulate at inner angle, with longer tcerebus setae
along inner edge.
munretsorP htiw eaxoc detacol a elttil dniheb eht ,elddim eht ecnatsid neewteb eht eaxocdna
lanigrambus suclus sa taerg sa taht neewteb siht suclus dna roiretna .nigram edisrednUdehtolc
htiw hsiwolley yerg tnebmucer mrofites.selacs
Length: 2.7-2.9 mm.
:epytoloH elamef epyT( .oN ,6692 uhsuyK ,).vinU ,tumaM ,habaS .E ,aisyalaM .41-11.iiiv
,1891 .K otomiroM.gel
:epytaraP 1 ,elam ,gniroP ,habaS .E ,aisyalaM .81-51 .iiiv ,1891 .K otomiroM.gel
Distribution. .E aisyalaM.)habaS(
Remarks. sihT si citsiretcarahc ni eht gniwollof :stniop murtsor ,rednels hcum regnol naht
,mutonorp eannetna ,mrofilif epacs ton gnihcaer ,seye tnorf eaibit etalugna ta eht xepa forenni
elgna dna artyle htiw owt esrevsnart sdnab fo mrofites.selacs
Biology. No information.
4. Endaeus robustus.von .ps
.giF()4
.elameF mreD hsiwolley ,nworb laretal snigram fo ,mutonorp mulletucs dna laremuh selgna
fo artyle krad nworb ot.hsikcalb
Forehead between eyes 213 semit sa ediw sa eht esab fo ,murtsor htiw a naidem .aevofseyE
ylthgils xevnoc morf .selpmet murtsoR a elttil retrohs naht ,mutonorp htiw a naidem dna owt
eanirac no hcae edis fo lasab ;flah 1annetna seborts gninnur yleuqilbo ot eht rewol trap fo.seye
Antennae inserted at the middle of rostrum; scape nearly as long as funicle, reaching eyes;
funicle with first segment robust, second 5/3 semit sa gnol sa ,tsrif driht ot htxis lauqebus ni
,htgnel ;esrevsnart bulc eciwt sa gnol sa ,daorb tsuj a elttil regnol naht dnoces ot htxisstnemges
of funicle combined.
Prothorax 1.3 times as wide as long, widest at basal one-third, constricted at the apex,
rounded laterally; dorsum coriaceous, with fine recumbent greyish setae. Scutellum -bus
,ralugnairt regnol naht .ediw artylE yldaorb ,etavo tuoba 52.1 semit sa gnol sa ,ediw -lellarapbus
sided at basal half; each interval with recumbent yellowish grey setae and medially a row of
subrecumbent longer dark or yellowish grey setae irregularly. Legs clothed with recumbent
70 H. KOJIMA & K. MORIMOTO
D
Fig. 4. Endaeus robustus .ps .von - ,A ;elameF ,B daeh d;nxaarohto r,pCtnorf
;gel ,D.annetna
taarom ee fhfhtotd ie whftl athuo bshaici h,whto ortalugna i hrat tiawrom etfno r;feat ehssiyerg
, hdr,tee d itttee yw auolvalel on trnaitt;tu eidnstictv bonlseeiriehtfodtpiw tcerebus egantoelsa
depahs-elk c,ipsr ah ht, six ewefl pogr ane dantenatin i,oe pgrdeenni .sucnu
dmn e ue h e te medteae,a ceewnih xaec anlttiwteoxlodeatsehhcodlnhttobetcdatsirbiilPmd
lani gsruacmlbuuss sa taerg sa tah tneewte bsiht suclus dn aroi r.e nteindgairsadrmeehdtnoUlc
.eates yerg hsiwolley tnebmucer htiw
:htgneL 8.3.mm
:epytol oeHlamef epyT( .oN ,7692 uhsuyK ,).v i,naUlhkgnoS .S ,dnaliahT .32 .iv ,5691.Y
ekat.agyeilM
Distribution. .dnaliahT
Remarks. sihT si tahwemos ralimis ot E. zonatus .ps .von yb eht tsubor ,ydob tubylisae
e lmb o asrdr rfeatetpit oecnensaih rteahh.tcyek
.ygol.oni oBoiNtamrofni
5. Endaeus formosanus .ps.von
.giF()5
.rethgil elttil a eannetna ,nworb ot nworb hsiwolley mreD .elameF
dae hneereowFte bseye l/2 ot 311 semit sa ediw sa eht esab y ,,lfxmtoeuhvrhgnttiosilcowsr
n.aai ed svexeom eyfm. vEusmn reoo rhtlrc,ecspfm tuom unrmRehtaottohhintswor panaidem
den aaon win troh ace1cadaei nsfl;noafesltaanbha seborts gynl iehnutnqauierlnbe obehtesab
NEW ASIAN ENDAEUS AND ENDAENIDIUS 71
d e.efamtonurnreetst snnaoi Are letrtoifl eeb hetlddi m;fmourtsor epacs gnihc a,esreyeregnol
n;a eheltlcciin nuhuftfi wt str niyeflm dg,aeeo strdabnvooces 3/2 sem is tagn o ,dlstrasirhoittf
htxis lauqebus n i,htgnel a e lrtettirl onh as;hdtno cbeus lyclr aeecniwt sa gnol sa ,daorb aelttil
ret r.noeahlhsctinuf
xar o5h3t.o1r-P52 .s1emit sa ediw sa ,g n,odlen e,tdteat layflligdno rdtesd nyulolraretaldna
nd , onrnhdii oice sheigilssehrrhluebteawi ortmft,upnsd ceaerhidd itpitlhiswaatwnc-oiecpnao
,ralugnairt bmuusllet u.ceSa tee lstanpebmu che trdiewht o,ldcerutc nyulpe nmiufs r;oeda thetsiw
a elttil regnol naht ,ediw htiw eni ftnebmuce relap .eates artyl E5.1-4.1 semit sa gnol sa,ediw
dedis-lellarap ta lasab ,flah tuohtiw lacipabus ;illac hcae lavretni htiw enif tnebmucerdna
de s.ntmsoxeaugpet iex pfydL e.ao liteelgna layetaaiP bepdwmseou mrc fero reeb gluantsotlil
dehtolc htiw tnebmucer elap ;eates tnorf aromef hcum rekciht naht eht ,sroiretsop htiwa
r a,lh uthgocnoiathiw rsti sse lnah tflah eh thtdiw fo e hatrome fta e ht ts;et dntiinewooaprifbit
y dltn lsie.auhhx njeteddrbpe eeavdttrnanualicid
munretsorP htiw eaxoc detacol dniheb eht ,elddim eht ecnatsid neewteb eht axoc dna
lanigram bsuusclus hcum retaerg naht taht neewteb siht suclus dna roiretn a.nigraemdisrednU
.ea tee lstanpebmu cee nrhi tfdiewhtolc
.elaM xarohtorP 52.1-02.1 semit sa ediw sa ,gnol tsediw a elttil dniheb eht .elddimartylE
.desopxe muidigyP .ediw sa gnol sa semit 6.1-5.1
:htgneL 2.3-4.2.mm
:epy teollaomHe fepyT( .oN ,869 2u h,sn ua,yh)K s.igvenipinimU ag,TnnaeY,iassHomroF
,,.,
2---
I/
0.
/’
/’
p.
A
Fig. 5. Endaeus formosanus ..vposn - , A;elame F,B ;elam ,C daeh, xdanraohtorp
.annetna ,F ;gel tnorf ,E ;elam ,ottid ,D ;elamef
