Table Of Content© ZooloIgnisctaiSltt u.tPee,t2 e0r 0s2burg,
StudoineN se arcstpiecoc Lfie eusc (oDpiisp Cthearmaa:e m yiidae).
IT.hree descorifpNteiarLocentu iccpo upbisl biesfh1oe9rd6e 5
VN..T anasijtshuk
TanasijVt.sN2h.0u 0kS2,t. u doinNe esa rscpteicocif le esu co(pDiisp Cthearmaa:e
myiiIdT.ah ere)e .d escorfiNpetaLireocuntc ipocup isb lbiesfoh1re9ed6Zo 5o.s ystematica
Ross1i1c(a11,9) 3:- 207.
ReviosfiNoenca t rycpoteLfise uc(opDiispC thearmaa,e mdyeisicdbraeiefob)1re 9ed 6 5
warse alTiyspeaednsc.d o rresmpaotnedwrieinrsagetl usd foirte hfodel loswpiencg ies:
L.ameriMcaalnla1o 9c2hL1,.a; t r!faAclider1si9 c2Lh5.,;fl aviAclodrrn1ii9cs1Lh 4.,;
maculTahtoam p1s9o01Ln; .m, aj oMra ll1o9c2hL1,.m; i nMoarl l1o9c2hL1,p.;a ra/le/a
Mall1o9c2hL1,.p; e mphMiagalel1 o9c2hL1,.p; i nipMearldla1o 9c2hL1,.s; i mplex
Loe1w8,6 9.
V.N.T anasijZtsohoulko,Ing sitciaRtluu stseAi,ca ande moyfS cienUcneisv,e rs itetskaya
nabI.S, t .Pet1e9rs9b0uR3rgu4 s,se i-am;ab irla:c h@z in.ru
Introd uction ofm ansyp ecuiseifesnm ga laerdseo omteobd e
uncerutnatsiionlmc el eadrelfiync erdi taer reia
Thsep ecoifLe esu cMoepiisg (eDni pCthearad:e velaonpided de ntwiifitpehad r tiscpuel caire s
maemyiwihdoalseae)r pvraeoeynA phiadna de whosmea lwea sc orredcetslcyrF irubtehde.r
Cocciadswa eeal,slP syl(lSitdeave1e 9n6s 7o;nm,o raem ,a lheolt oy wpiet fehm aplaetr yap (oe rs
McLea1n9,9 8p)l,aa yv eriym porrtoalin net v icvee rqsuai)ot fetb eenl otnodg i ffesrpeent
manbyi oceHnoowseesvste.ur do ifte hgsir so upci es.
havbee eunn eviendn i fferreegnitAo lnmso .s tIi ntteopn udb lasi esrhoi fae rst iocnNl eeasr c
70s pecoifte hsgi esn aurske n owfrno mt h etiLce uctohpoiinssie d;s e vottsoep de cwihei sc,h
Palaeabrulctet tsihsca 2,n0 s pecwieerdsee werpeu blibsehfoertdeh w eor ko fM cAlp ine
scrifrboemtd h Nee ar(cMtciAcl p1i9n6 e5,,( 19p6u5b)l iisn"h Ace adt aolfto hDgei ptoe fra
197179,7T 8a;n asi1j9t9sn6ho)aut,lko l,fw hi chAmerniocraot fMh e xicToh"id.sa tweac sh o
currebnetllotynot g h giesn Tuhsi.cs ab ne e xsebne caduessec ripputbiloilnsasht weeedr r e
plaibnyte hdfae c tth daett ermionfma ot sitotny piccaarlrloiyuie tndm ordee twaiitdlhr aw
specoifLe esu cboype ixst emrnoarplh ol ogiincogafstl hmea lgee ni talia.
featuirsae lsm oismtp osbseicbalouefts he e ir
uniforTmhietrye.afo d rees,c roifop ntseip oen Matearnimdae lt h ods
ciceasbn es imitlota eornt h(eo rfcs o utrhse er e
arseo msep ecwiietdshi stifenacttuitrvhe eas t Durimnysg t ianyt hUeS Ai n1 99I2v ,i s ited
areea styod etermsiomn oess)tp, e cciaebns e thceo lleocftt hiMeou ns eoufmC ompar ative
deteromnilbnyyep dr epaorfta htmeia olgnee niZooloHgayr,vU anridv e(rMsCiZtw)yh, e Ir e
talSiiam.id leatre rmiunsaifetnmigao lntee rstudtihteeyd op fLe e ucsoipmipLsol eeWwxh. i le
minailnsio wate dlelv elaottph meeod m ebnut t,i nC anaidn1a 9 9I4s ,t udtihveead cs otl le ction
hapso tenatsdi eamlo,n sbtyrR aats(ep1di9 85o,fL euciontp hiCesa nadNiaatniC oonlalle ction
198169,9 109,9 6R)a,s& p Bie rto(l1i9n 9i3o )f,lnsOetctta(swC,aN CD)rJ;.. M Fc.A lpgian vee
Bescho(v1s9k9ai5n )Bd,e scho&v Msekr iz meh idsr amfta nuscarbioLpuettus c ionpcilsu d
(199H8o)w.e avfe emra leev,ei cnfo llaenc dteidnt ghs ep echiehe assst udTiheedms.ae n us cripts
reaornet dhs ea maep haisds sp ecX,i ceasbn e inclauk deeuyds ienxgt emrnoarlp hoblu otgy,
lontgos pecYi;te hsee x cepatrimeoo nnso withgoeunti sttarluiccT thueIrnw e o.r kfoer d
phagsopuesc sipeesc,ci leesad relfiyn aobn let hrmeoen twhistt hhc eo lleocfLt eiuocniosn p is
extearnpaple araansndpc eec,ri eeasir nce udl thNea tiMounsaelou fNm a tuHriasltW oarsyh,
tufrroem k nowpna rebnymt est hdoedssc riibnegd(tNoMnN Ha)nt dh Iel liNnaotiuHsri aslt ory
byG aim&aT ruirn (e1r9 9S7od),e. t ermin aStuirovnUesryb,a( nIaN HISr )e.c eailvases ode r ies
194 VN.T anasijNtesahrusckpt:ei ccoi fle esu coIp• iZ sO.O SYSRTO.S SIVoClA1. 1
oft yspep ecifrmoemDn rsW. .P ulawCsakliiw,i thhoeuatwd i)t,th hs ea mlea b"e4l65 6A8c"c.o rding
fornAicaa deomfSy c ie(nCcAeSsB) e.s idteors e coart!d NsH Sl,a b"e4l6 5m6e8a""n Dsi pdteera
thessoemm,ea tewreirsaetl usdi itneh cdeo llesctroaypihnigd s; IEvlMalan.ys 1 t,51o 9n1,D2 .. K .
McMil lan".
tioofnC SIR(OC anbAeursrtara,an ltdih ae)
DescrFiepmtaiBlooendl..ye n2g.t1hm- m2 ..2
ZooloIgnisctoaifltt huRetu es sAicaand oe mfy
Bodlyi gghrteH ye.a1 d. 5t-i1m.ae6shs i ag sh
ScieSntcP.ee st,e r(sZbIuINrnt)g o. It s atlu dileodnFgr.o2 n.s7 -t2i.mn8ea sr rtohwhaeenr a d.
morteh 2a0n0 s0p eci mens.
Ocelpllaawrte ea kmlayr kfreodn;st tarldi a prek
Aftreert utrnoSi tn.gP etIer rescbeuair vge,d
grewyi,tt whdo a brrko swtnr oinip tesssi O dT e.
signipfiocrat(nimtoo tnrh ea7 n0s 0p ecimens)
ind1e.xL6 u.n umleed ium-Asnitzeaendndn .a
oft hceo lleocLfte iuocnbosep liosnt gCoiN nC ga ribsltaa3c rkad,r isttohmrteeiremel e osn ger
anNdM NHp,r ovitdhfiern agm eowfto�i r skt han 2nhde.i 4gGt heitnm laee sts hsea yne
stutdoby eg reaextplayn AdsDe rdM..c Alpme
heigMhitn.ub trei satrlreasin,nog nesedp a rse
(perscoonmamlu niacsastuiimoatenp d)p, e ars
rowp,r esoengn etn aan;t ebrriiolsrat rlgee st.
ratehxetre nsive.
Lateraolfm esstornigopotelusdbm ir soehwx n,
McAlp(i1n9ee6 n5u) me1r5as tpeedco i fes
tendtiodn igs 1t/ao4lfm esonoMteudmi.a n
Leuc(ot phaies snu bgoeLfne uusc otphwiaest r) e strgirpeAeyrs.e b aest wseterindp eevsoo bfiri ds
descrfriobmte hdNe e aracntodin cfre o mt h e
tloevs2e /ro3 fm esonTowtpoua mi.or fds o rso
nortNheeornt r(oCpuibFcais)ov .fteh e(mb elclean btrriaspltr leesLsee ngbtsl. aa cpki;oc fefes
Loe1w8,6 b6e,l lWuillal i1s8t8bo9in,v, i tmtoartaaanb da soetfs i byiealelo onfow rl;ee tgasr si
Mall1o9c4ho0,c, e lMlaalrli1os9c 4ah0n,v d e rneaernltyib rleaolcnymk i,d hainnldde fi grss t
ticMaallilso1 c9h4w,0e )r sei ngoluet d btywtoa rsoymeelrlOeonsww i.n vgesi,Mn asn d
McAlp(i1n9e6i 0ntt)ho se e paorcaetlel aRr icsonviednri gse1t /at3l1p, . t3i mleetssh sa n
groTuhpe.gi ern eric caormbebe imngad 4ti+i5s sotenacslto ifCo uLn.a rbgleas cpkco otv enng
chanbgyTe anda sij&tG sahium(kaip rnri e ts os)
synte1r+gu2ip tt ope o s?tree drgitew;ro ?a etr
thgee nusLeuacloowpniig1tn 5nha e ,sw p e cisemsa.slplo ts opntr eers3ge;1dn taterm ke dian
Ouotft hree mai1n1oi,nn (ego rbMiatlall oicsshp ,optrse soenan ntt eerdeigo oftr e rg4i-t5e.s
192i1as)s ynoonfyL m.a rgentZiectotlelrTiasx onnoomtTiehcsde .e scroifMp atliloonc h
ste1d8t4(,8M cAl&p Tiannea si1j9t7sI2h )u.k ,
(1 92i1is)n disFtoeirxn acmthp.ewl reo,"t Teh o
havset udtihteeyd pa enosdt hmeart eorfti h ael
rawxi otwrhi thtowbuort o vwint tbuaated " d,e d
10r emaisnpienctgih reees s;bu elittnshsg eu b-
thtahtte y psiecra"ilhe tashst e h ovriatx'.·tI ante'
jeocftt h sitsu dy.
reatlhipetr ye,s oeran bcseeo nflc aets etrrailp es
Unfortunantoaetllo lyft , ht ey psep eci mens
ivse sreyr tiaoxuosn cohmaircaa ncLdta. em re,r i
wermea lweh;ai mtso r-seo metsipmeecsi mens
cansae nMsaul lmoacbyhec onceaiascv oemd
that wearsme a licenit theeddie rs cr iptions
ploefsx p ecSiteusd.ty hiteny gsp eer diined os t
turnoeudtt bo efe malTehe.r erfoerdee,s c ription
soltvhpeer ob-laelpmla ratwyeprfeeem sa le,
ofs evteyrpailens se ictohmeprnl oesrta et isfac
soa naloyfms ailgsee niitisam lpioas sible.
tory.
Neatrht ey psee riinIe NsH iSsa t uwbei th
Malgee niwtearsleti uadi ginle ydc derraiwn;
lab"e4l6 5(6t8sh"ae ma est ypaent)dh see c ond
inwgesrm ea diegn l ycjeerlTilemnyri.n �oyl o-lab"eLle ucaompeirsiM caalnla<o lcJeh. t,R. .
accorMdciAnlgp(1 i 9n)8eO.1 T (o cetlnlaanr
MalloIcctoh n"t.5a m ianleaedns1 d 4fe malIe s.
glei)n dreexfet rots h rea toifto hd ei stbaence
prepaaltrlhe emdsa ela ensddi scotvhteahrtee yd
twepeons toecreialonltrdih d ei stbaentcw ee en
beltoLon. gg lyp hiniTvaonraas i(jTtasnhauski j
anpyo stoecreilaolnrtud h sae n teorn ieo.r
tsh1u9k51 89,8 I6it)ts .h erperfoorbeta h balte
thtey sppee ciamlebsneosl toont ghs ip�sie e s,
Descriptions anLd.g lyphinisvhoobrueaal cdc eapastJ eudn
iosry noonfyL m.a meriBcuattnh aei.rns eo
Leucoapmiesr iMcaanlal1 o9c 2h1, distbianstciots porrro evjteeh ctiths e sis.
DistriIblultiIintfooh nsii.pss e. c iiisde esn ti
Mall1o9c2h31,5: ;41 94207:2 . cawli Lt.hg lyphi-nneiavcroolrsyam opolitan.
Modeo fl ifAet.l eaisnpt a rlta,rfe veaode n
Typperse seirn/v NeHdHS o.l otsyppeec i(mwehni cah phiItdfh ssi.ps e ciiisde esn wtiiLtc.gha l ly phi
wagsl uoenpd a pterri alnogseltme,p)p t iyln a be"l#l ed
nivoirtlasa, r feveadoe n n eaornlheyu ndred
46568""L,e ucopis Maaml-eltr.yi pc(eaw"nr aii tnt en
penc"iTly)Lp,ee u coapmiesr iMcaalnlado "cr'he,ld a specoifae psh ids.
_
be"lI NHTySp e#0 122O"n.t hsea mpem a rteh r ee
puJaarniadsa m atlulcb oen tatihnaeib ndgo imngeln y cLeucoaptirsi faAclider1si9 c 2h5,
eriTnh.ia sb do(mweint dhoouufrbtot m,h olostpyepcei (Fi1g-s5 )
meni)fes m alneom,ta laesM alltohcohu ght.
Paraty1p esc;,l:,a b"e4l6s5 6"8la"l,o tlyepuec opis
ameriMcaalnla<;>o "c",h! NHTSy p0e1#2 32" ;(<;> o neA ldr1i9c2h15,5: 2 .
ZOOSYSRTO.SC SAIV o1l1.• V N. TanasijNtsehaurksc:pt eiccoi fLe esu coIp is. 19 5
2
4
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Fgis1 -L5e.u csoa ptirifaAcliders1i ,2c ,h-.h 3e,ma adl;ge e niitnap lrioafi4 l,ee p;a ndfrroimub me hi5n,ad p;eo xf
aedeafrgoumbs e loSwc.a0 l.em1:m .
TypeHso.l ot<;>y,pp ree,s eirNnv MeNdHL .a be"lTsy:p eS ta1t2ed' :,7 <;> ,G oldenAduag2l.0e1 95e9m A.p r.
#2 82U1S2N M"",S aFnr ancCiaslci"of.E "W.,a lthe1r9 6p0r,e Add.e lpgiecse( aRe.M Gi.t chIed' l,2l <;> ),;
col"lI.cn"o ,n newcittThiheo cno dippilnois-irsa ditahsteaa mepe"l ,aacnecd o lle"ccto8ol rl2-,.0- 59"C.a li
"LeucaotpriifsaA ciidPe"as.r atNyMpNeHs6:: (c;>" alforniaI:cl ,S aFnr ancJias1nc.9o 2,o8 n,A delsgp e.s
lotyrpeeap"la,lr ya atny5dp" ep aratwyipttehhss ea" m)e onP in(uEsW. a lth1e r(9)? B;)a ,l dLw.i6 n5 0S0.' B,.
labtehlCesa ;l ifoArnciaad eomfSy c ie(nCcAeSs4) c;> ,, Co.1,3V 195(5W .RM.aMs.o Inc;> ),A;l baAnlya,m eda
thsea mlea bels. Co1 5V I19 5a8d,u clotl leocnMt oendt eprien(yeF r ed
Othmeart eerxiaamli CnAeSd3:d'. , 2 c;> ,S aFnr anD.BenneIt<;> t,t) hs;e a mlea bbeulVt,I 1I9 538d' ;,3
cisCcaolI 2. 7 2.5 p,r ebsyE. .W altchoel(rlo o.nn d'e 9,t hsea mpel aacnecd o llePciutnsro ard,i VaI1t 9a5,8 ;
penlcaibl"e Lle ucboep/iOls.a S .a?n"ad n otlhaebre l2 <j> ,S aRna faeMla rCion1, 9 V I1 95p8r,e odnP. i neus
"LeucaotpriifsaA ciidDe.es tF .. BDe.n n1e9t5t8a "n)d, o nM onteprien(yea slple cidmeetnesr mbiyFn .eDd.
I'i' ,t hsea mlea bbeulXt,1 81.9 2C6N.C W:a shi ngtBoennnaesLt .at t rifaMceixeisIc)<jl o.,3: m iE E lS aito,
19 6 V.NTa.n asijNtesahrusckpt:ei ccoi feLse ucIo• pZ iOsO.S YSRTO.S SIVoClA.1 1
DgoM ex8.2 0J0u'l,4y ,1 96(4J .MFc.A lpi5n9 e,1) 0; Leucfloapviisc Aolrdnri1is9c 1h4,
miW ElS aitDog,o .9,0 00J'u n5e- 161 96 4(Fi6g-s 8 )
(J.F.McAZlIpN2i:c! n ,Ae l)b;a Cnayfl,A.i l ameCdo a
VI1 95o8nP inruasd i(aFtraDe .Bd e nne(trte)c eived
fromD rJ.. MFc.A lpiinen xec hanCgSeI)Rl.Oc! :, 1 Aldr1i9c1h44,0: 4 .
9,G oldGeantP ekS .FC.a,l ifoFreneidaoi,nnP ign eus
spb.r,e Jdu n1e9 3(8F .GJa.y ). Typesp reseirnNMv NeHd. H olotd'y.pL ea,b e ls:
DescriMapltefei,mo anl.Be o.dl ye n1g.t7h- "BrownTsevxiM.la ly1e 0 1,9 10""R;AV .i ckCeorly
lect"o#5r "4;r;" e lda b"eTly Npoe1 84U2.8S .N .M.";
2.m2m .B odgyr eHye.a( dF i1g-s2o )fc! 1. 5
relda b"eHlo loLteyupceo jlpaivsi coArlndirsi ch''.
timaeshs i gahsl onogf9, 1 .(3a verAangtee).
Paratlyd' p,5e 9 s,t: h sea mpel atciema,en c do llector.
ripoaror ffrt o nasn guilpnar ro fiFlreob.na sr elCyNCl:9 ,t hsea mpel atciema,en c do llector.
wideannetde irnic! a,2d .;1 -t2i.mn2ea srr o werDescriMapltefei,mo anl.Beo .dl ye n2g.t0h-
thhaena idn9, ,2 . 2-t2i.mO4er sb.is ttralilip gehs2t .m6m .B odlyi gghrteH ye.a1 d. 7-t2i.ma0e s s
grewyi,d eannetde Orciealpdll.aa ertl eo ntgraithei agshl onFgr.os nosm ewwhiadte annitneg r iad;
angusllairge,hl telvyFa rtoensdtt.ar alil plseio g h2t.6-t2i.mn7ea sr rtohwahener a Fdr.ou nssu ally
greFyr.ow nismt ahn tyh wihni htaeie rssp,e c imaolnloytgorneheyo ;w evmearpy,o sstewsdosa rk
nealru nuFlaecl.eu ,n aunlpdea orftg endaa r kg,resyt riflpoewsia nrgo uoncde lpllaaartn ed
neabrllayLc ukn.us lheod rotr,bs oarlcd uerr v erdeaclhuinnuWgl heenfr. o nmso notoocnee l,l ar
sligOhTti lnyd1.e. xA6 n.t ebnlnaa3c rkad,r istpolabtaer oeultyl iinnse pde;c iwmietdnhas r k
mer4e-t 5i mleosn tghea2rnn dG.e nhai g2h.,4 -fronsttarli appepseo,ab rlso ng-tOrcie alnlgiular.
2.t6i mheisg thheearny h ee igMhitn.ub trei stulneesvsepnalfrcyoe mde acoht hOeTri ,nd veaxr y
ong enaarr anignre odwa ;n tebrriiolsrat rl gee isntbg,e tw1e.ea5nn 2 d. S0c.a apnepd e dilci eglh t
oftseunrr oubnyds emdab lrli sPtallbeplsia. c kb.rowbna,s oflagealnlador miyesertleal roaw;
Mesonowtiutmhs oturtic poevse,br uentdo , t tiboe twleeennog ft3 hra dn 2dn adr issetgal
densbeylr ye,g ulloanrg itruodwoisfsn maal l lmenvtasr frioemd1 .2t:o21 . 5L:u1n.uh lie g h,
briessTt.wl op aiorfds o rsocbernits rtadlloe rsesdaglse e mi-roGuennhdaee id4g..h 5t - 5.0
presFeenmtob.rl aaw cikty he llaopwi cteisb ;i ateimleests hsea ynh ee igShetv.eb rrailsfo trl..me s
blawcikty he llboaws aelst,h osuogmhe t imienisgrr egruolpwar re soenen atcg he naan;t e rior
femoraan tdi beinatei brlealccyok l;o roa ftiborni sltalregPsea sldtpa.irb kr owGnr.em ye d ial
tarvsair iea.bgel.ne,t, i brlealocyrfik r,ts atr sostrsilpiegsdh atrltkyhel arin gg hrteb ya ckg round
meroenms i ddalnhedi nldeb glsa acnksd l ig hptrleysoenmn ets onottougmew tihtwehir d lea t
yellboaws aolrfil ryts,at r soymeelrloeonsaw l lersatlr oifps eism illiagrchlotyl oreaxttieonnd
legOsnw. i ngvse,iM n asn Rd4, 5 sligchotnlyi nbge yomnidd odlfme e sonoAtruebmae.st w een
verigdnei s1t/oa3rla rpea ratlvple erslyh. oi r ntm edisatlr idpeevsoo ifbd r isatrlebeaes t;w een
compawriidtsihos nst eaclto ifCo urn;a ti2is.o 0 -medainaldla tsetrrawilip tsehms a blrli sotn lleys
2.A2b.d omsiynnatle l+r 2ga iltmeoe sntt i reexltyentdoai nntge 1r/iTw3oo.r p aiorfds o rso
browtnhc;io sl orsaptrietoatonde sr 3gb iytt we o centbrrailsp trleessFe enmto.br laa acpki, c es
waveRsa.t hweirdm ee dilaonn gitsutderi inayaell ltoiwb;yi ealel moiwd,do lfae n tetrii boira
presoentn etr g3i-to5er 4s - 5D.a rfike lod fs darketnaeyrdes;li l eoxwc,ae ppittc aarls o meres
modimfiiecdr otarbiscfrehonimlta a et eedrga elsd arkeOnnwe idn.g vse,iM n asn Rd par allel
4+5
offi rasbtd omtienragilincl t . es ors omewchoantv enregaairpn igc tep1s. .1 -1.3
Malgee ni(tFail3gi-sa5E )p.a ndmraisusm i vtei,msehso rtthedarin s ptoarlto ifCo uAn.b domi
extenidnivtneogrs yh ocrutr,ve eddiH tyepsa.n nasly nte1+r 2dg airtkeae nnteedr tiwosorm l ayl;l
drinuema relryeA cetd.e algounswg i,d e nebdlaocrgk o ldsipsohst osm eteivmiedsoe nnt
apictaulrnlueypd,G. o nopaobdssep natr,a m etreers3gs,ii tmeit sloam ram leld sipaolot nts e r gites
larsgmeo,o tthalpye red. 35-.T hecoafme i nubtrei sotnal bedso minal
TaxonnoomtIiencgs e.n isttarlut chtseup reet ergeintceisrb cysl meabdlu lnt o ticbelaabclke
ciiessc loosnelt yoL .p inipMealrdlaoI ct hss.p oDtasr.fik e lodfms o dimfiiecdr otornichiae
determbiyne axttieaornpnap le airssai nmc pel lea;teedrgaoelffis r asbtd omtienragolifm t aelse s
coloroaftt hiheoe nai dnc ombinwaittaihbo nn oetv ident.
senocfse t roinpt ehmsee sonaortveue mrc yh arMalgee ni(tFai6lg-isa8E )p.a ndmreiduimu m
acterBiuistitt ns ie cc.e stsocao rnyst ihdpeeo rs sizweidt8,h- 1p2a iorfls a rbgrei satftleneros ;
sibiolfai m tiys tianck oem parwiistdohan r ktabcloen streixctteiinondnmts,eo d ium -sized,
facesdp eciomfLe p.ni sn ipfreordmwa h,i Lc.h ratthheeirdn i Htyepsa.n dwriiduemin nae ndt e
atrifdaicffioeenrslis ytn h set ruocftt humera e l eripoarrs tl,i gchutrlvAyee dd.e awgiudsei nne d
geni talia. baspaalsr tm,o ottahpleytr oiwntaghr sidln i, g htly
DistriUbSuA(t Wiaosnh.i Cnagltiofon rn,i rao)u,nadpee dx .
Mexi co. TaxonnoomtiLejlc.sa .v iccoarbnned i est er
Modeo lfif Lea.r vparee daotnAo dresls gp eps.m ineeads biylt yhy ee llcoowl oofut rhb ea so
onP in us. flagellTohmiecsrh ea.r aicsut neirqa umeo ng
ZOOSYRSOTS.CS AIVo l1.1 • V. N. TanasijNtesahrcustkpi:ec c oiLfe esu coI pis. 197
Fig6s-L 8e.u copisjlAalvdirc6iom,cr ahnlgi.ees n iitpnar loifi7aal, ee d;e fraogmbu esl 8o,w ;g onaonppdao rdas m eres
froma boSvcea.0l .em1:m .
stuNdeiaerdsc pteioccLfi e eusc oOpnti hcse.o nothPearlbspl.ia L cakt.se trraoilfp meess onotum
trasrtyr,u ocfgt eunrieit nsao lctih aa rac tienrciosntsipocirhc aurocduolsny s pimceudo iuasl,
andde termoinnolayntt hiciohsna rcaacbnt e ers trfaiipnentsol tyi coenanlbeyala nert eerdi goer.
unreliable. Mesonsoutrafaelcv ee nly bcymo ivne urteed
DistriTbhusitpsie ocinknis.oe wson n frloy m brisatrlebeaesst, ws eterni poeust lTwnioon te d.
Tex as. paiorfds o rsobcrenitsFrtealmleob srl.aaw cikt h
Modeof lifuen known. yelalpoiwtc iebsai;nat dea yr eslilO onww i.n gs,
Leucompaicsu lTahtoam p1s9o1n0, vtehdaiinnas snpRtMdo4+:a rp ltoa 1frC0 autOnl.1pn l a . eb1.ldt-4;oi1 ( mm sFeehi9nsog) r.,t er
(Fi9g-s1 3) synteI+r 2ga iltmeeon stti brreolwryno ;u n ded
paisrpeodpt rse soentn etr g3i-ts5eo;sm et imes
Thomp1s9o1n20,3: 8 . simillaarorgnlte ye rg3i-mt4ie,ns ou ntt ee r gite
5,b uuts uaelvledyni lmyi npiosshtieOnrng i ad.
'J:yppreess eirNMNHnv .e d Holot9y,"p Cea,lN veebrt. t erg3i-t5e,s t brliaacnskgp uoaltlaspsr ro e sent
10/15"/9,0 Ty9p/e"1#,3 14P1a"r.a tI9y ,pt ehssea: m em ediLaalclkydi.an firgek l odmfso dimfiiecdr o
labIecl l,t; h sea mlea bpellau,ds d itliaob"neBalrl e d
tricohnei dageoe ffis r asbtd omtienragil in tes
fromEricoopleolrtaid(seIt nnhds eeis s"c. rTihpotmipon,
sownr ottheta ytsp eer wiae3ss a cln 1d )9. mal es.
Othmeart eerxiaamli CnNeCdC:.a naAdlab:e2 rtcl a,: Male g(eFnii1gt0sa- l1Ei3pa)a .n drium bear
Scan9dV iI1aI9, 5 (60P .e cMka)n.i tI9o ,Wb ian:n ipieng1 g0p aiorfbs r isotnpl oessts eirdpieao,sr s ing
Sept1.9 2e7m,. 131.9I2Xe7 x,g rassacsl (eA . V.smooitnhdtlioys traolulneydd eidHt yepsa.n d
MitchCe.nHCe.ur rr,a 4ncl ),5;9 ,W pg1.8 .2X7g ,r assr iusmh omrats,s piavsesfr,io nmbg e neianttho
coc(cpirdo Ab.aMVbi.lt yc h4ecl n,2e9 r,W) p;gF .e b .29,
paramcelroesfuses lewydi itthg; o nocpoomds
192(8A .MVi.t chNeMnNeHrI:)n .d i5acl n,1a 9 :,L a
FayeVtIt-e9(,-J 1.A5Ml .d rich). plerteedluyc ed.s lAiegbdheetnsaltmgy,ou ost h
DescriMpatlifeoe,nm .aB loed.ly e n2g.t2h- lyt apetroiwnaagpr edx .
2.mm8. B odlyi gghrteH ye.a1 d.t 6i maeshs i gh TaxononmoitcTe hsse.p ecieisae ssdi ilsyt in
asl onFgr.os nosm ewdhialtaa ntteedr iadg;u iasviheenarx btleearn paple abryta hnpecr ee s ence
agi2n.gt6 i mneasr rotwhaenr Ohrebaidt.oa fpl a isrpeoodtn ts e rg3i-wt5ie,ats lh m coosmt
strliipggehrset Oy c.e lpllaaortu et loinnble yyd pleatbes eonflc aet setrraoilfpm eess on otum.
ligghretcyo lorpartoicoetneo,dw ealdru dnsu lGee nisttarulciotcfLu .mr aec uliuasnt ac ommon
asn arrtorwi aFnrgolnsett.ar glir peey , dfoarLr ekuecrog poinso;pc oodmsp lreetdeu lcye d.
coloFurroedndes.n sceolvyew riectdoh l o urlLe.msasculhaaatsnaa lmcoosmtp laentaelloy
haiOrTsi .n dex1 .nA5en.at re annnadar i sgtoaucso untientrp haPera tl aeraerpcrteisce nted
bla3crakdr, i st5to immeaerlsseo a n2sgn Gde. n abyL .sz epliget1i9i3sA7ofc, arz ree lp,ofr rtoemd
hei3g.h1t- 4.l0e tsthsiea mynhee es i gBhrti.st hree gfrioomHn u ngtaoMr oyn go(lTiaan asij
tloengs e nvaa riiannbu lmeba errr,a in1ng- e3dt sh1u9k8,I6i t)ds .i stinfrgouLmi.m sa hceu dl ata
irrergouwalsna;tr e rior britsh talnient hnpeor teasboelflny ac telse atrrraogilfemp ree sso notum
19 8 VN.T anasijNtesahrusckpt:ei cocif Le esu coIp• iZ sO.O SYSRTO.SS I CVoAl 1. 1
/
12
Fig9s- 1L3e.u cmoapciusTl ahtopams o9na,. b dom1e0mn,a; lg ee niitnpa rloifi1al1 ae,e; d eianpg ruosfi 1l2ae,e; d eagus
fromb elo1w3p,;a ramfreormbe e low.0 .Sm1cm a.l e:
(t hohuigghvh alryyii ningn teoncfso oilrt ayt ioDni)s,t ribCuatnia(odAnal. b eMratnai,t oba),
narrgoewneaarn, id nd ifferiengn ecneist aUlSiA(c N ebrIansdki aa,n a).
stru-cptaurrtei ncaurlrasoruwlreysr,lt oyb leasrM odoelf if Lea.r avrapeer edoanEt roirosp eltis
anade deagus. Hoowpfea viesrrpe,od t spscr ae lseesn.ce
ona bdomtienrag3li-a t5ne sdsi mibleatrwieteyn
basgiecn icthaalriac(c mtaesrsrsio vuen,d Leedu copisM amlalj1oo9c r2h 1,
sursltoyblgeaesrn, e raolfa esdheaaapgbeu s(,F i1g41s-6 )
senocfge o nopaoridens d)i coafact liovrseee
latiobnesthwtiehpee snsp ee cTiherese .l ation
Mall1o9ch23,15 :2 .
shbiept wteheesnsp ee cmiaeaysl b seoj udged
byt hfaec tth tahtel iarrp vraeoeyns cafrloems Typesp reserivnINe HdS H.o loty'i'p.Lae b,e "lSst:. Joseph,
cerepaalrtsi,co unsl paeroclEfiyr e iso pEevlitIilsMla..3y ', 1 4""S;aF lotr" kLe"u;c ompaisjM oarlT lype." (in
dentthleyss,pe e chiaevoser igirnealtaetdip veenc"liTylyL )pe;eu cmoapjisMo arl l<;,o"(c rehld a be"lNH)S;
recefrnotmalc yo mmaonnc easntsdoe rp,a raTtype e#1 d2 4P"ara type3s'i' :,th esa mel abOenlpe.ar aitypse
greasoenwdeit, h ohuetadA .l lomtyepnet iboyMan leldao bch
duteog eograipshoil caatli on.
sePnrto.b iastbe lrvfoyer pd r epaoragfte inointa lia.
ZOOSYSRTO.S SIVCoA1l 1.• V. N. TanasijNetasrhcustkpi:ec c oi/eLucseo pIi s. 199
16
Fig1s4 -L1e6u.c opmiajsoM ra lhl.1o 4ch,e a1d5;a, b dom1e6mn,a; l gee ni(taaflMtiaealr l och).
DescriFpetmiaBoloned.l.y e n2g.t4hm- m2.. 6 TaxononmoitcMe sa.l l(ooccphi. tw .r)o te
Bodlyi gghrteH ye.a( dF i1g41.). 4t-i1m.ae 6ss " tyfepmea lael,l oatnyfodpu erfe, m alpea ra
hiagshl onFgr.od niss twiindcetanlneytd e ritaydp,easnp"dr, e setnhsteke edt dcrhaywo ifn g
2.2t-i2m.ne6as rr otwheharen a Hde.as dm oothmlaylg ee ni(tFai1lg6iA.)ac . c ortdtoih dnirgsa w
rounidnpe rdo fiFlreow.ni stm ha nlyi grhetl,ai nLg.m, aj ohra asr atmhaesrs eipvaen drium,
tivleolhnyag i presr,p entdofri ocnuOslr.ab ri tmaolderwaitdeeedl iyat c eusr,hv yepda nda rium,
strilpieggshr tes yl,i gwhitdleyan netde rilaagdrea. edeaapgoadle amneadr, a tshmearl l
Frontaild esnttlriiicwgpahheletv ln iy e frwoe md aedeaBgututh sdi.rs a wiiisnn ge xtahpcert e;p a
fronbtu,fat rd arwkheervn i ewfreodm aborvaetu.is ofoenrdt hdirsa wwiannsgo fo tu nidtn h e
Ocelpllaanrto reta idsiesdt,i nognyulb i yshIeNdHc So llectiotnhm.eo seBtxe psriedsessi,v e
ligchotl orAanttieoornci.eo lsrla umssei az se charafocrtt hseepr esca irteehl sei gfrhotnw si th
posteornieOosTr.i nd1e.x4 -L1un.u6la.er cthhiani rresl,a twiivdgeeel nayan t dh rpeaei rs
shapdeodr;ps aawrlit t sha mheas ia rsfr on so,fde (rautnhceorm cmhoanr aMcatlelerov;c e hn
proclAinntaebtnlena.aac r ki,s tbar od3warnrd;kw roatbeo "utth arnesdeo metfoiumrde osr so
arist4o.m0e-tr4ie.m a5esl s o nags2 ndG.e nac entrUanlfosr"t)u.an maotNneegla yr,Lc etuicc
hei3g.h3tt- i3m.le4ets sh sea ynhe e iMgihntu.t oep isse ebnym eI,h avneofo tu nmdo rsep eci
bristlesi no narerer gaurnlogaowern dg enam;e nwsi stuhca ch o mbinoafct hiaornaa cntde rs
somaen tebrriiosfortrm lientsgi cglhuts wti etrhc,o unlomdta kper eparoafmt ailgoeen nsi talia.
onlea rbgreirPs atllbpeli.aL cakt.es rtaroli f pesD istriIblultiinooni. s.
mesongootludmi sh bronwanr, r orwMe,oldaoetfl i ifuvene klnyo wn.
extentdopoi sntge praioiofrdr eG .r emye d ial
strriaptephsae lerex ,t entdmoii dnodgfl mee so
LeucompiinsMo ar ll1o9c 2h1,
notuamr;eb aest wseternui spueadsle lvyoo fid
(Fi1g7s- 24)
briosnta lnetse1 r/oi4fo mre soTnhroetpeua mi.r s
ofd orsocbernitsprtralelesF see nmtob.rl aa ck;
Mall1o9c2h31,5: 41 ,9 4207:4 .
tibbilaaweci kty he lalpoiwcfr eesq,u eynetlly
loown i nnseurr fatcaery;se il ltowwao,p ical
Holotyd'p,pe r,e seirvn!e NdH LSa.b e"lDsBu:o iIsl,l .
tarsodmaerrkeeOsnnw eidn.vg esiM,na snR d4+5 Aug9.,' 17""L;e ucmoipnMisoa rlT yple."( ipne nc"iTlyp)e;
conviednri gs1et/ at2p1l; . 3-t1i.ml6ee5tss h s a nLeucmoipnisMo arl ld'o"c( hr e"dI)N;THyp Se #0 12 5".
dissteaclto ifCo unO. n a bdom(eFni1 g5 .) ,DescriMaplteiB.oo ndl.ye n1g.tm5hm .E n
synt1e rga+il t2mee o nsttid rae(rlgkyo lwdhiesnht ibroed gyr eaistecsdo ,l, oa ucrcdoirntg o
viewfreodma boavned bfrloambc ekh i nMda)l.lloecahd,-H gera(edFy i1.g7 1.). t 8i mae ss
Sligohbtllipyqa uiserp eoodtnt s e r3gc iotleo urheidagl sho nFgr.on nowsti deannetde 3rt iiamde,s
identPiacmlaeeld lilyao.nn gisttuprdriieans aelnn atr rtohwahener a Odr.b iatnfradol n sttarli pes
ont er3g;ibt reim gehdti satnrp iraeeso e nnetq uadlalrSykh .ot rhtbi rni sptrleesosnefr not n s
terg4ai nt5de. s anddo rpsaaorlft l uOncuellpell.aac rta er d iform,
200 VN.Ta nasijtNesahruckspt:ei cci ofe Lse ucoIp• iZ sO.O SYSRTO.S SIVoClA1. 1
2 0
., :-··.·. :- : ·
t
21
·...... .........
I
18
23
2 4
Fig1s7 -L2e4u.c ompiinMsoa rl lo1c7hh,e. a 1d8;1, 9 m,a lgee niitnpa rloifiaaln fredo mb el(oawft MecrA lp2i0n,e );
aedeaagnaudps o deimnpe r ofi2l1ea,;e deafrgoumbs e lo2w2s;,u rsltoyblieanpsr r ofi2l3es,;u rsltoyblfreao srm
behi2n4gd,;o nopaondpdsa ramfreormbe esl Socwa.0l .em1:m .
ZOOSYSRTO.S SIVCoA/l! . IV. N•.Ta nasijtNsehaurksc:pt eiccif leesu cooI pis. 201
sliglhitfaltryeo dua nhdt eorcieolOrlT ui sn.d exTaxononmoitcDe est.e rmionfta htsiipsoe n
1.8O.c ellii nse iqzuLeau.ln ualrec -s hacpiebedysd. amatgyepdse p eciwmietsnhh, od ret
Antenna3 rabdrl iasctk6o;t m iemaresles o nags scriopftM iaolnla oncpdha rdtelsyt rgoeyneid
2n.dG enhae i5g.ht2ti mleests hsae nyh ee ightta.li isva e ruyn relAimaobnlmgea .t erfrioaml s
Minubtrei sptrleesosen gn etn iar,r egaurlaCralnyaa dnaUd S Aa rsee vesrpaelcw iietsshi mi
rangoendle,a rgPeaslbtpl.ia Nceki.t lhaetre rlaaglre nitbautlthi eahy,a vlea taenrmdae ld ial
nomre dsitarlvi ipseoisnbg lree amseesdo notsutmr;oi npt ehmsee sono tum.
Malldoicndho d ti sctuhcsihssa raicnht ideser DistriIblulti inooni.s .
scripMtciAolnpw;ih nsoea ,tw h tey sppee cimeMno dofe l ifuen kno wn.
befoprree paorgfa etniiowtnra ol(tiileani , tt ht a.t)
"mesonwoittuhdmoo urts ocsetnrtiTwrpoae ls "L.e ucpoapriasMl allell1oa9c 2h1,
paiorfds o rsocbernitsprtraleles sFe enmto .r (aFi2g52s,6 )
blawcikty he llaopwi cteisbb;il aaewc ikty he l
lobwa sfoerse;t aresnit ibrlealficyrk s,1t - 2 Mallo1c9h23,15: 3 .
tarsoymeelrleoosnmw i ddalnehd i ntda rsi.
Holotypc;>e,p, r eseirn!v NeHdLS a.b e"lMsu:n cIil el,.
Wincgrsu mpvleeindno,dst i sceArncicbolred.
Ju5i,' 14""S;t oCnry." "L;e ucpoapriaslM laelllao ch,
intgoM allotchship,sie echsa "st hsep ootns
typ(eip"ne nc"iTly)Lp;ee u cpoapriasl Mlaelllaoc;> "c h,
secotnedr gsimtaaeln lwd i desleyp artahtee (dr,e" d!)N;HT Sy p#e 012 8".
postmearrigooirffin r tsetr ggriaptyue b es cenDte"scriFpetmiaBoleon.d.I y7 . 5mm lonlge ,a d
(actually ptohiean rtsresye nfetre1er+ng2ci egt ree Hye.a( dF i2g51.). t4i maeshs i gahsl ong.
antde r3g) i.te Frosnlsi gwhitdleaynn etde 3rt iiamndea,sr r ower
Malgee nitIra elcieafri.ov mDe rdJ. . MFc.A lthahne aOdr.b isttarlli ipgeghsrt eo yne xte rior
pisnkee tdcrhayw oiftn hggese niotfta hlteiy ap ebordaelrosen ygme a rgdianrg;kr enye atroe r
specimenb,yh imiman 1d 9e6( 8F i1g8s- 19m)i.ddolffre o nwsh,e trheec iorl moeurrg weist h
Accortdoti hnegcs eer,rc ait lhaerrhgy ep,a ndfronsttarlBi opteoh.r baint(dat llo e sesxetr e nt)
riusmh acpoem mtoomn a nLye uc-ocpuirsv efrdo nsttarlci opveesbr ype edr penddaitrchkui lna r
ipno st1e/rP3ir.oe rp aorfga etneio tvaelrical berairsthlaeiOsrc seo.lrp l laanrto eetl evoantleyd ,
edn,o hwa rpdelryc epDtoripsbaalorlefet . p andsliglhitglthyht afrenor n sttarlOi Tpi en.d1 e.x 5 .
riucme,r acnihd,y panddriisusmoa llvm eodsL tunuwlieda er,c -shwaipttehhd bi;rn i sot rles
entiarneldlo tysh tes ihrao pnela;ye dewaig tuhsh aiArnst.e annnadar ibsltaa3c rkad,r istomere
apod(eFmie2g 02s,1) s ,u rsltoybl(eaFsri2 g2s, 3.t6i maesls o nags2 nDdi.a moeftbears oflagel
23g)o,n opaonpddrasa mer(eFsi2 g4w.)e rwee lllomreartelh aer2rg1. te i,m leests hsea ynhe e i ght.
preseSrvuerds.lt oyblteahsiri pn nr oafinlfrdeo m Gena h3e.it6gi hmtle ess s thheain ghety.e
reaGro.n opaondpdsa ramoefer qeusla eln gtMhi.n ubtrei satrlreasin,nrg e egdur loawpr,r esent
Aedeatghuissnl ,i gehxtplayni dnbe adsp aalr to.n g enaan;t ebrriiossrlt ilgelh otnltgyhe parrn e -
Fig2s5 -2L6e.u copis
para/Mlael/lao 2c5h ,.
hea2d6a;,b do men.
202 V.N.T anasijNetasrchtuikc: osfpL eecuiceoIsp• iZ sO.O SYSRTO.S SIVCoA1l 1.
cedionngeP sa.l bplia Mceks.o nowtiutmh oduetlto(iHd.BeEis.s d3ed' e,)C ;a rryPilnaPgcr ei nce
stricpoevseb,ry er da tshpearrr osweos fb risEduaCroAd.u g2a. n 4d,1 96e1xP emphgiagl(ulNs.s H .
AndersId'o ,tn h)se;a mpel aJcuen1,e2 1 ,9 6e2xP, e m
tleTsw.op aiorfds o rsocbernitsprtraleles se nt.
phigguasl( lHs.B Ei.s dIe9 e,S) u;l t7Ia In6I,8P ,o pu/u;
Femobrlaa acpki,sc leisg yhetlllyo twiibsihab;ae ls amif(e?rQ)au. e bIed' c,:I 'i' ,M acdoCnoallpld. ,
alssloi gyhetlllyoa watip sihcfi ertssa;tr so meJru1ie9 s7P3e,m phgiagolunpls o p(lEa.rAHl.l eyUnSeA ):.
yelldoiwso,tn aedlsa rOknw. i nvgesi,Mn asn d MontaId'n ,aI 'i': , 2m iW. B iFgo rVkI2,I1 .1(9B6.7A .
R paratlp1l .et5li ,ml eess sd itshstaeancl t iFoonotOeh)iI.od' :,I 9 ,1 2m iN EA lliVaIn1.c8 e1.,9 70
1+5 (T.SPl.u sAsr)i.z 2o n'i'a,S: o utRhewsS..t aC.o,c hise
ofC uL.a rggoel d-brsopwponrtie ssohena n bt
County1V6I-H1-(9B6.9FA o.o tNeM)N.HU :S AD:. C.:
dominal 1+ t2ae nr3dg i(tF2ei6s g) ..
Id' ,I 'i' ,C olumFbeidpa.a ,or n.P empshpt..r ansversa
TaxonomicT hnseop teecisivs.ee src yl otsoe l iesx,F eb2.6 /(7E9. SAa.f veIrnrdyi)1a. 'i'n a,L: a
L.simpLloeextw h;se o dlies tiinnac ptpieoanFr ayeItV2t 6e( ,J .AMl.d riLcohu)i.s3 id' a,Pn tAa l:l en,
anciets h ger eastioezfret hbea soflagellSeopm1te5-. 1r 5e( T..JHo.n eId's ,)B ;a tRoonu gMea,y1 3.
DistriIblultiinooni.s . 16i,s sMuaeyd1 4- 16b,r ewdi tPhe mphpiogpuusl i
trave(rTs.JuHos.n e2s'i' ),t; hs ea mpel aAcper1,.1- 1 8,
Modofe l ifuen known.
issAupe2rd8. 1- 8b,r ewdiP. tp hop ulic(aTu.JlHoi.ns e s);
ld' ," Feedoin5n 4gg.a olnlE lm-leaf'.
Leucpoepmipsh Miaglaleo1 c9l h2, DescriMpatlfeiemo,an l.Be o.d lye n2g.t0h-
(Fi2g72s,8 ) 2 7.m m.B odlyi gghrteH ye.a1 d. 5-t1i.ma6es s
higashl onFgr.os nosm ewwhiadte annitnegr iad;
Mall1o9c2h31,5: 0 . 2.8-t3i.mn0ea sr rtohwhaener a Odr.b isttarli pes
ligghrtefr yo,n tsatlrv iaprey iincn ogl oration
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