Table Of ContentOdonatologica33(4):433-436 December1,2004
SHORT COMMUNICATIONS
Heliogomphuschaoi spec. nov.,
a newdragonfly fromsouthernVietnam
(Anisoptera: Gomphidae)
H.Karube
KanagawaPrefecturalMuseumofNaturalHistory,499Iryuda,Odawara,
Kanagawa,250-0031,Japan
Received 11November2003 /Revised andAccepted20May2004
Thenew sp.is described, illustrated andcomparedwith thesimilarH.selysii Fraser.
Holotype<3:SVietnam,Lamdongprov., 15kmfromBaoLoktoHo-chi-minh,6-V-1997;
depositedinKanagawaPrefecturalMuseumofNaturalHistory,Odawara,Japan.
INTRODUCTION
During myrecent survey ofSouthernVietnameseOdonata,manyinteresting species
havebeendiscovered.Theregional faunaispeculiar forthehighaveragenumberofen-
demics,associatedwithmountainstreams. KARUBE(2000,2001)andHAMAlAINEN
& KARUBE (2001a, 2001b) brought onrecordalready somenewinteresting species
fromthisarea.Here,anewHeliogomphus isdescribed.
HELIOGOMPHUS CHAOI SP. NOV.
Figures1-14
Material.—HolotypeS'.southernVietnam,LamDongprov., 15kmfromBaoLoktoHo-chi-minh,
6-V-1997,depositedinKanagawaPrefecturalMuseumofNaturalHistory,Odawara, Japan.— Paratypes:
samelocality asholotype,1 9.5-V-1997; - 2S, 12, 16-VI-1996;- 63.32,3I-V/1-VI-2002;- 2(5,
6-V1-2002. All H.Karubeleg.
Etymology, - Dedicated tothe memoryofthe latedoyenofChineseodonatology.ProfessorDr
Chao Hsiu-fu.
MALE. — Head black withgreenish yellowmarkings; baseofmandibleandgena
palegreen;labiumwhitishyellowwiththeapexbrown;labrumdarkbrownwithapair
434 H.Karube
ofdorsaldrop-shaped markings whichis thinattheinnerpartandbroadendlaterally;
anteclypeus brown;postclypeus black;antefronswithabroadbandatthecrest;vertex
notdeveloped; occiput black, ratherroundly raised.
Thorax. — Prothoraxblack, withapairofyellowspotsnearthelateralapices, with
apairof smallspotsadjacent eachotheratthemiddorsalpartofmedianlobe.
Pterothoraxblack,withsymmetricallygreenishyellow marksas follows:antehumeral
stripetaperedtoventrad,withjoiningneartheendofmesothoraciccollar;mesokatepister-
Figs 1-14.Heliogomphuschaoisp.nov.(male;Figs 1,2,4,6-11;female;Figs3,5, 12-14):(1)headinfron-
talview; — (2-3)thorax, dorsal view; — (4-5) abdomen, obliquelateralview; — (6)caudalappendages,
lateralview; — (7)same, dorsalview; —(8)accessory genitalia,lateralview; — (9)same,ventralview; —
(10)penis,lateralview; — (11)distalsegmentofpenis,ventralview; —(12)posteriorpartofhead,oblique
frontalview; — (13)same,frontalview; — (14)apicalsegments,ventralview.
Heliogomphuschaoisp.n. 435
numwithatriangularventralspot;mesepimeronwithamedianstripe whichstopsclose
toposterior margin; metepistemum withbandtaperedtodorsad; metepimeronlargely
greenishexceptformetapleural suture;poststemum greenish yellow;outsidesofcoxae
yellowish. Legs ratherlong, black.
Wings hyaline; nodalindex: forewings with 13-16antenodalsand9-12postnodals,
hindwings with10-15antenodalsand 10-13postnodals; pterostigmabrown,expanded
atthemiddle,2.2-2.Sand2.5-3.0mminlengthinfore-andinhindwing, respectively.
Abdomen.— Black withgreenishyellow markings; 1stsegmentwith abandat
ventralhalf,withthintriangularmarkings atthedorso-posterior area;athinlongitudi-
nallineruns onthedorsalcarinafromtheanteriorborderofthe2ndtotheendofthe
7thsegment;segment2 withlateralmarks attheauricleandwithdimventralband,the
latteris connectedwithmedio-posterior markingnearborder;3rdto8thsegmentswith
asmallspotonbase,9 to 10thsegmententirely black.
Caudalappendages darkbrownandslender.Superiorappendage darkbrownexcept
forawhitishapical half, forcipate with theapex strongly curledupwards, with strong
lateralspines atthebasal 1/5andalsowithbroadlatero-ventralprojections atthebasal
half.Inferiorappendages very widely divaricated,thelobetapered apicad andrather
curvedupwards, apex withadorsalspine.
Genitalia. — Anteriorhamuleinvisibleinlateralview,strongly inclinedbackwards
andcrossedinnerpartof posterior hamule,slenderandlong, thinnerfromthemiddle,
stronglyhookedbackwards attheapex;posteriorhamuliratherbroad,bill-shaped,with
apex triangularly pointedanteriad.Vesicle developed,expanded dorso-anteriorly,with
ashallowmediannotchattheanteriorarea;stemslightly bent;middlesegmentrobust,
tapered toapex;distalsegmenttrumpet-shaped; posterior lobesemicircularly protrud-
ed, withsmalltubercullarapex.
Measurements (mm).— Abdomen(incl.app.)31.7-34.1,hindwing26.7-28.5 inlength.
FEMALE. — Maculatepatternsimilartothoseofmale.Greenishyellow markingsof
labrummoredeveloped; vertexwithstrongspines atthesmallridgebehindeach lateral
ocellus, andwithapairofvestigial spots attheposterior slope ofvertex.
Wingshyaline;nodalindex: forewingswith 14-16antenodalsand 10-13postnodals,
hindwingswith11-12antenodalsand 10-13postnodals; pterostigmabrown,expandedat
the middle,2.9-3.0and3.0-3.3mm inlength inthe fore-andhindwing, respectively.
Yellow markings ofabdomenmore developed, athin longitudinal lineruns on the
dorsalcarinafromtheanteriorborderof the 1sttotheendofthe7thsegment; 1stseg-
mentwithbandattheventral2/3;2ndto3rdwithmedio-lateralband;4thto7thseg-
mentswithawedge-shaped spotoneachbase; posterior partof4thwithvestigial one;
8thwith dimlypatch; 9-10thsegmententirely black.
Valvular valvae semicircular, with the apex notchedand bilobed, with the apical
partofeach loberounded.Cerci darkbrownand about2 timesas long as 10th, with
pointed apex.
Measurements (mm). — Abdomen(incl.app.)34.5-34.9,hindwing31.0-31.6inlength.
DISTRIBUTION. - SouthernVietnam.
436 H.Karube
DIFFERENTIAL DIAGNOSIS. — Twospecies ofthe genusHeliogomphus havebeen
hithertorecordedfromVietnam,viz.H. scorpio andH. retroflexus(ASAHINA, 1996;
TSUDA,2000).
ThisnewspeciesissimilartoH.selysidescribedfromLeito,Burma,andalsorecorded
fromAssam(FRASER, 1934)andThailand(ASAH1NA, 1981,1986;HÄMÄLÄINEN
&P1NRATANA,1999).Itiseasily distinguished fromthelatterasfollows: (1) latero-
ventralprojections atnearthemiddleofmalesuperiorappendages, (2) femalevalvular
valvaearebroadwithroundedapex(valvular valvaeofH. selysislenderwithtriangu-
larapex), (3) femalewithoutapairofpost-occipital spines, (4) femalewithapairof
spinesbehindthelateralocellus.
ACKNOWLEDGEMENTS
IwishtoexpressmydeepgratitudetoDrMAMORU OWADAoftheNational ScienceMuseum,Tokyo,
who gavemeanopportunitytocollect inthearea.IalsothankMr ITSUROKAWASHIMA forhissplen-
didillustrationsofthethorax andabdomen usedinthispaper, andtoDrYUTAKAARITA,DrSYUHEI
NOMURA,DrHIROTSUGUONO, andDrSATOSHISHIMONAGAfortheirkindhelpinthefield work.
— Apartofthisstudyissupportedby theGrant-in-aid No. 13575015 forField ResearchoftheMonbuka-
gakushoInternationalScientificResearchPrograms,Japan.
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