Glyptosceloides dentatus, a genus and species of Eumolpinae new to Chile (Coleoptera: Chrysomelidae)

8 Pages·1994·3.1 MB·

Rev. ChilenaEnt.
1994,27:69-76
GLYPTOSCELOIDES DENTATUS, A GENUS AND SPECIES OF EUMOLPINAE NEW TO
CHILE (COLEÓPTERA: CHRYSOMELIDAE)
INGOLF S. ASKEVOLd' AND R. WILLS FLOWERS"^
ABSTRACT
Study of known South American genera described in the tribe Eumolpini, sections Edusites and
Iphimeites strongly suggests that specimens of a previously unidentified species of Eumolpinae
represent an undescribed genus and species. The genus Glyptosceloides is described as new, from
Chile, includingthenew single species. G. dentatus.
RESUMEN.
El estudio de los géneros descritos enla tribu Eumolpini, secciones Edusites e Iphimeites, indicaque
los especímenes de una especie no identificada hasta ahora representan un género y especie no
descritos. Sedescribe unnuevogéneroparaChile, Glyptosceloides, con unasolaespecie G. dentatus,
n. sp.
INTRODUCTION cies were described, rendering study of the
Neotropical Eumolpinae a difficult task. It is
The Eumolpinae of Chile comprise 11 or 12 difficult to rationalize description of new gen-
genera, including about 27 described species era in this context. Numerous taxonomic prob-
(Askevold, in prep. a &b). At least 5 of these lems persist, well illustrated by the genus
genera are endemic to this narrow country, Jansonius Baly (Askevold & LeSage, 1990).
which is bounded by desert to the north, ex- ISA has examined the type specimens of all
treme cold to the south, and the Andes to the species of Chrysomelidae described from
east. Two of these genera, Stenomela Erich- Chile, and RWF is familiar with almost all
son and Hornius Fairtnaire, are perhaps the New World genera ofEumolpinae. Therefore,
most primitive of the subfamily, making the despite existing taxonomic confusión among
fauna a rather unique and limited one. Chil- Neotropical eumolpines, we are confident that
ean species of genera that are not endemic to the genus and species described herein must
Chile are generally single members of groups be distinct from all other Neotropical Eu-
that are widespread in the Neotropics, some molpinae, readily recognizeable among Chil-
even extending into North America. ean and other Neotropical genera.
Over the years that Jan Bechyné published Terminology of female genitalia follows
on Eumolpinae (for references, see Seeno et Lindroth (1957); that for male genitalia fol-
al., 1976), a large number of genera and spe- lows Askevold & LeSage (1990).
GLYPTOSCELOIDES, Askevold and Flowers,
new genus.
Entomology-Biological Control.
Plant-lnsect Interactions, DivisiónofAgricultural Sci- Type species: Glyptosceloides dentatus
ences, Florida A & M University, Tallahassee, FL 32307-
Askevold and Flowers.
2926, USA.
(Received: November 3, 1993. Accepted: February 18,
1994)
70 Rev. ChilenaEnt. 21, 1994
Etymology: The species is named in reference
to the broad profemoral tooth.
DESCRIPTION:
mm
Length, pronotum and elytra 4.50-6.00
mm
((ScS), 5.00-6.80 (99); width at humeri
mm mm
2.30-3.00 ((ScS), 2.80-3.60 (99).
Body (Fig. 1,2) obovate; head, pronotum,
elytra, undérside bronzish, bronzish-black or
metallic green; femora and tibiae entirely palé
reddish to entirely bronzish-black or metallic
green, tarsi dark with metallic sheen, in most
specimens individual antennomeres, tar-
someres, femora and tibiae reddish at extreme
base.
Figures 1-2: Habitus photograps ofG. dentatus: (1) dorsal
aspect, (2)lateralaspect.
Head with labrum apically emarginate, with 4
or 5 subapical setae placed in emargination.
Etymology: the genus is named in reference Frons and clypeus densely, coarsely punctate,
to the genus Glyptoscelis Dejean, the Chilean punctures confluent in some specimens to
species of which, G. pulvinosus (Blanchard), form longitudinal rugae laterally and on vér-
G. dentatus resembles in size, colour and tex; surface between punctures coarsely mi-
elytral rugosity. croreticulate; clypeus emarginate apically,
with setae at outer angle ofemargination; each
Diagnosis and Description: pygidum with in- puncture of frons with short white prostrate
complete, broad, shallow median furrow, the seta. Eyes elongate-oval, weakly emarginate
furrow flat (Fig. 10); profemur with large next to antennal insertion.
acute ventral tooth at midlength (Fig. 5); tib-
iae multicarinate, the carinae prominent but Antennae with scape irregularly globose;
incomplete; tibiae strongly flared at apex; pedicel subglobose, slightly shorter than
prothorax with distinct ocular lobes, lobes scape, distinctly shorter than segment 3; seg-
projecting ventrally and forming discontinuity ments 1-4 reddish brown, remaining segments
with prosternum (Figs. 3, 6); prothorax, head blue-black, all segments with scattered white
and mandibles of male enlarged. Because the setae; 3-6 elongate, slightly wider at ápices; 7-
genus is monobasic, description ofG. dentatus 10 flattened, asymmetrical and subserrate, dis-
provides other specific structural details. tinctly wider at ápices, 7 and 8 broader than
following segments; 11 narrower, spindle-
GLYPTOSCELOIDES DENTATUS shaped, apically conical.
Askevold and Flowers, new species.
(Figs. 1- 17) Mouthparts. Maxillary palp with apical seg-
Type specimens: Holotype d (Museo Na- ment spindle-shaped. Mandibles large, robust
(larger in (ScS), laterally uniformly curved,
cional de Historia Natural, Santiago): "Hun-
apical teeth broad, slightly acute, with broad
garian Soil-Zool. Exp. CHILE: Prov.
occlusal edge; with outer surface coarsely
Coquimbo, Socos, 12.XI.1965 / Nr. P-B. 113
densely punctate and with long white prostrate
leg. Mahunka". Of the remaining 142 speci-
setae.
mens examined, listed below, 113 are desig-
nated paratypes.
Prothorax (Fig. 3,6) subquadrate, L/W = 0.62
Type Locality: Socos, in the province of Co- - 0.76 (dcS), 0.56 - 0.70 (99); disc medially
finely punctate, punctures separated by several
quimbo.
Askevold&Flowers: NewgenusandspeciesofEumolpinaefromChile. 71
'PLD
OL-
Figures 3-11: Structures ofG. dentatus: (3) prosternum (arrowis pointofmeasuringwidth); (4) claw; (5) leftprofemur; (6)
prothorax ofc?; (7) left9 mesotarsus; (8) left (S mesotarsus; (9) apical two abdominal sternites ofcf; (10) pygidium ofd";
(11) tegmen of aedeagus. Abbreviations: AD = apical declivity; CXi = procoxal cavity; EPMi = proepimeron; EPSi =
proepisternum; MF=medianfurrow; OL=ocularlobe; PLDprosternal-ocularlobediscontinuity.
72 Rev. ChilenaEnt. 21, 1994
times their diameters, laterally with mixture of fourth; protibiae somewhat curved, wider in
punctulae and very coarse punctures, sepa- apical half; tibiae multicarinate, slightly tc^
rated by approximately their own diameters; moderately sulcate between carinae, and with
surface between punctures coarsely mi- long setae arranged linearly in sulci, meso-
croreticulate to shagreened in some speci- and metatibiae somewhat flattened on anterior
mens, especially laterally, disc laterally surface. Tarsi densely and uniformly pilóse
somewhat swollen, obscuring lateral margin beneath; basal tarsomere of all legs expanded
((S(S), or not swollen (99). Lateral margins (especially (S(S), those of fore- and middle
with distinct uniform bead, evenly curved legs (Fig. 8) more strongly so (almost as wide
(dorsal view), directed strongly obliquely as long) than those on hind legs; second tar-
downward from posterior to anterior angles; somere broadly triangular, with acute apicolat-
anterior angle directed forward; basal margin eral angles; third tarsomere shorter and
with uniformly broad, flattened marginal bead, narrower than second, deeply bilobed, with
bead with single row of shallow punctures; lobes rather slender. Claws robust, with broad,
each angle with single erect short seta in large basal, slightly acute tooth (Fig. 4).
puncture, anterolateral seta situated sublater-
ally on anterior bead. Proepisternum with an- Elytra coarsely, irregularly punctate on disc;
terior margin convex, forming ocular lobes, sparsely to densely, coarsely transversely ru-
ventrally distinctly raised and discontinuous góse, especially laterally and at midlength;
with anterior margin of prosternum; surface most punctures separated by distance equal to
coarsely punctate, each puncture with long or slightly greater than their diameters in most
seta. Proepimeron glabrous, coarsely punctate, specimens; surface between punctures mi-
punctures separated by distance greater than croreticulate, punctures toward apex becoming
their diameter, intervening surface mi- regular, forming striae; intervals convex to
croreticulate to shagreened. Prosternum (Fig. costate, with regular, single row of punctulae,
3) anteriorly recessed for reception of base of punctulae in apical área bearing very short
head; anterior margin prominently raised as seta; humeri prominent, rounded; basal calli
slender high margin, surface markedly excá- weakly developed; postbasal depression ab-
vate behind margin; entire surface with long sent. Sides subparallel, slightly converging;
slender erect setae; broad, 1.10 X diameter of apex conjointly broadly rounded. Basal mar-
procoxa, widened behind coxae; longer than gin moderately to markedly costate, costa
either mesosternum or metasternum. obliterated toward scutellum. Epipleuron nar-
row, acutely raised, slanting inward, tapering
Mesosternum flat, broad, slightly narrower evenly from base to apex, finely setose api-
than prostemum, width = 0.80 X width ofmid- cally.
dle coxa, densely and moderately coarsely
punctate, with moderately long prostrate white Scutellum triangular, base 1.40 X length; with
setae. Metasternum short, slightly swollen punctures absent to several, fine to coarse.
anterior to hind coxae, only slightly longer
than mesosternum; densely and moderately Abdomen with basal segment slightly longer,
coarsely punctate, with moderately long pros- remaining segments subequal in length; sur-
trate white setae. face densely and moderately coarsely punc-
tate, with moderately long prostrate white
Legs (Fig. 4,5,7,8) robust, densely punctate setae, surface between punctures finely alu-
and sparsely covered with long prostrate setae, taceous or microreticulate. Apical sternum
surface alutaceous. Femora strongly swollen with shallow subapical median depression in
in middle, entirely palé reddish brown to en- both sexes, apical margin shallowly emargi-
tirely infúscate or with only extreme base red- nate medially in both sexes; male with median
dish; profemora with large ventral tooth at subapical área glabrous, microreticulate, im-
mid-length (Fig. 5). Tibiae gradually widening punctate, reddish brown (Fig. 9); female with
toward apex and strongly expanded in apical similar but smaller median subapical área; 4th
Askevold&Flowers: NewgenusandspeciesofEumolpinaefromChile. 73
SBF
AM
14
ALS
Figures 12-17: GenitaliaofG. dentatus: MALE: (12) apical view ofmedian lobe; (13) lateral view ofmedian lobe; (14) in-
ternal sac sclerite (dorsal view). FEMALE: (15) ventral aspect ofsegment VIII, hemisternites and gonocoxae; (16) dorsal
aspect of hemisternites and gonocoxae; (17) spermatheca. Abbreviations: ALS apicolateral setae of hemisternites; AM =
apicomedian setae ofhemisternites; AS apical setaeofgonocoxa; A8 - apódeme ofsternum VIII; B =baculum; BH -basal
hood; BLS = basolateral setae ofgonoxa; BMS = basomedian setae ofgonocoxa; BS = basal spur; CS = coxostylus; GC =
gonocoxae; HS - hemisternites ofsegment XI; IR =internal lateralrod; OS = orificial sclerite; PP = dorsal portion ofhem-
isternites of segment XI; SBF = subbasal fenestra; SD - spermathecal duct (about halfits length illustrated); SG = sper-
mathecalgland; SOS =suborificial sclerites;TG=tegmen.
74 Rev. ChilenaEnt. 21, 1994
segment with broad lateral depression, extend- gous and merely use the term internal rods for
ing onto 3rd segment in some specimens. convenience. Spermatheca (Fig. 17) with
elongate, transverse meshes; slightly con-
Pygidium (Fig. 10) with longitudinal median stricted at ramus; spermathecal duct attached
groove broad, shallow, flat, becoming obsolete basally, very elongate, forming few loóse ir-
on apical one-fourth; its lateral margin slightly regular coils (only half illustrated).
acutely projecting inward; with subbasal bis-
inuate line, and subapical biarcuate line de- Intraspecifíc variatíon. Specimens are metal-
marcating slight declivity, apical declivous lie green or bronze; legs vary in color from en-
área coarsely punctate, smooth between pune- tirely reddish to entirely metallic; punctures of
tures; apical margin crenulate; lateral margin the pronotum vary in size, and the intervening
carinate. surface various from distinctly microreticulate
to coarsely shagreened; punctures of the head
Sexual dimorphism. Female like male except vary in density and the extent to which they
as follows: head smaller, mandibles smaller coalesce to form rugae.
and less robust; pronotal disc laterally not so
swollen, the lateral margin entirely visible Distribution. Specimens have been examined
from above; basal tarsomeres of all legs not only from the Coquimbo Región, Chile. No
expanded (as in Fig. 7); apical sternum me- specimens have been found collected outside
dially punctate and microreticulate, not so ofChile.
deeply impressed.
Material Examined. Chile. Región de Co-
Genitalia and associated structures. MALE. quimbo: Prov. Elqui: Depto. Ovalle, Tres
(Fig. 11-13) Median lobe (Fig. 13) in general Cruces EP Reed (CAS 3), 4-XII-37 EP Reed
shape typical of Eumolpinae; with long, (CAS 1); Cuesta S. Cruz [may be Santa Cruz
lightly scierotized basal hood, lacking distinct mineral, near Tres Cruces] 1938 Reed (USNM
apodemes; with subbasal fenestra; with promi- 2; CAS 2); La Serena l.Nov.l957 LE Peña
nent basal spurs; tegmen robust, fused along (CMNH 1); Coquimbo XI-61 (OHIO 2; ISAC
midline (Fig. 11); internal sac with pair of 1); Vicuña 15.X.1983 L. Peña (ISAC 1), no
slender suborificial sclerites (Fig. 12) and ba- date Reed leg. (USNM 4); S[an]. Isidro (CHIL
sal irregularly shaped sclerite complex (Fig. 1); Dept. La Serena, Las Cardas Sept.
14; apical when endophallus inflated); flagel- Nov.1947 EP Reed (CAS 1), 3 December 1949
lum extremely long, rather abruptly thickened EP Reed (CAS 12). Prov. Limarí: Tr[anque].
at base. FEMALE. (Fig. 15-17) Sternum Recoleta, R. Hurtado 2.Nov. 1957 LE Peña
VIII with short basal apódeme; sternum other- (CMNH 1; ISAC 1); Socos 12.X1. 1965 Ma-
wise only with poorly scierotized apicolateral hunka P-B Nr. 113 (ISAC 6; RWF 4; TMB 54;
área; apical margin with numerous sparse ex- CHIL 3); Rio Guatulame 13-XI-1961 LE Peña
tremely long setae; hemisternites with cluster (BMNH 1); Manquehua, Majada Blanca 1.64
of apical setae; paraprocts separated into pair (OHIO 5; ISAC 2; SMC 1); San P. de Quille,
of slender dorsal rods with cluster of apicome- S.W. Punitaqui 15.Nov.l961 LE Peña (CMNH
dian setae; baculum distinct, apical, almost as 2; ISAC 1). Prov. not determined: Incienso
long as gonoxa; gonocoxae large, with apical [=Flourensia thurifera] El Pepe 7-XI-44
ring of setae, with subapical scierotized cylin- (CHIL 1;CAS 1);
der and basolateral unsclerotized área, with
cluster of 6-7 basolateral setae, with cluster of Additíonal specimens found in press, not
6-7 basomedian setae; coxostyli distinct, with seen by authors:
several distinct long setae. In addition to these Prov. Elqui: Totoralillo 14.X.91 E Arias
structures that are normally found in the fe- (CHIL 1); Las Cardas, 40 km S la Serena
male external genitalia, there is also a pair of 4.XI.52 R Wagenknecht, sobre Flourensia
very slender, oblique, internal rods (Fig. 15); thurifera ("incienso" (CHIL 22); 746 [hand-
we do not know with what these are homolo- written Philippi label], 1089 Myochrous
Askevold&Flowers:NewgenusandspeciesofEumolpinaefromChile. 75
flourensiae / Ph. 1209. Coq[uimbo]. [hand- punctate, surface laterally not shagreened be-
written Philippi label] (CHIL 3). Prov. Li- tween punctures; and larger in size.
man': Socos 9.XI.91 T Fichet (CHIL 1); El
Divisadero (E Punitaqui) 23.XL1971 G Rojas Vianeta. Four species were compared with G.
(CHIL 1). Prov. Choapa: Conchalí (N Los dentatus: V. varians Bechyné, V. argentinen-
Vilos) 22.VIII.1984 G. Carrasco (CHIL 1). sis Bechyné, V. uberaba Bechyné and V. po-
doxenus Bechyné. They are similar to in the
Comparison with other genera. toothed profemur; proepisterna forming ocular
Glyptosceloides was compared with various lobes; raised anterior prosternal margin; and
other Neotropical genera of Eumolpinae. Eu- transversely rugóse elytra. Vianeta differs in
rysarcus Lefévre, Vianeta Bechyné and the complete and uniformly deep pygidial fur-
Prionodera Chevrolat bear considerable re- row; legs rufous in most specimens, basal tar-
semblance, but are evidently not congeneric. someres not so expanded in males; mandibles
Few genera of Neotropical eumolpines have not enlarged in males; tibiae not carínate ñor
both enlarged mandibles and toothed pro- sulcate ñor widening toward the apex; profe-
femora, therefore Glyptosceloides is unlikely mur not so robust; and more elongate, slender
to be confused with other genera. body shape.
Argoa. Argoa species compared were A. rugu- Prionodera. Four species were compared with
losa Lefévre and A. tibialis Chapuis. Of all G. dentatus: P. bicolor (Olivier), P. lútea
Neotropical eumolpine genera known to us, IZ,richson, P. kirschi (Lefévre) and P. mar-
Glyptosceloides appears most closely related shalli Lefévre. This genus shares with Glyp-
to Argoa, with which it shares the following tosceloides the toothed profemur and raised
characters: 1) toothed profemora; 2) lateral ru- anterior margin of the prosternum. Prionodera
gosities on elytra; 3) incomplete median differs in the complete lack of median furrow
groove on pygidium; 4) sexual dimorphism in of pygidium; proepisterna not forming ocular
enlargement of hind basitarsus; 5) sexual di- lobes; pronotum, head and mandibles not en-
morphism in shape of pronotum. Glyp- larged in males; basal tarsomere only weakly
tosceloides can be distinguished from Argoa expanded in males, distinctly longer than
by the following combination of characters: 1) wide; and larger size and more elongate body
body shape obovate (oval in Argoa); 2) mandi- shape.
bles of males much larger than those of fe- Among Chilean eumolpine genera, only
males (no sexual dimorphism in mandibles of Dictyneis Baly (of authors) has toothed
Argoa); 3) apical half of antennae subserrate femora; however, this genus belongs to the
(submoniliform in Argoa). Myochroites and is distinct in pygidial shape,
toothed lateral pronotal margin, vestiture
Eurysarcus. Four species were compared with (scales) of the prothorax and elytra, tubercu-
G. dentatus: E. abiroides Lefévre, E. dejeani late elytra and all femora toothed. Like Glyp-
Lefévre, E.foveicollis Lefévre and E. rufinus toyceloides dentatus, one species of
Lefévre. They are similar to, in shape of the Glyptoscelis in Chile has similarly trans-
pronotum; proepisterna forming ocular lobes; versely rugóse elytra; Glyptoscelis species,
markedly broadened basal pro- and mesotar- however, have scales and setae covering the
somere of males; enlarged mandibles of entire beetle, lack'the femoral tooth, and a less
males; and broad and flat basal pronotal mar- well developed (to almost absent) pygidial fur-
gin. They differ from Glyptosceloides in lack row.
of profemoral tooth; pygidial furrow deeper
and complete to the apical margin; elytra not Placement of Glyptosceloides wíthin Eu-
to hardly transversely rugóse, with indistinct molpinae.
to quite well developed costae laterally and on Reid (1993:61) noted that "at present it is im-
declivity; tibiae enlarging apically but not so possible to define any suprageneric group of
markedly costate; basal pronotal margin not the Eumolpinae to include all of its currently
76 Rev. ChilenaEnt. 21, 1994
included members." Similarly, it is not possi- pest, Hungary (TMB); Dr. David H.
ble to ascertain which genera one should ex- Kavanaugh, Department of Entomology, Cali-
amine to determine the most appropriate fornia Academy of Sciences, San Francisco,
position of a new genus within the existing CA 94118 (CAS); Dr. Richard E. White, Sys-
classification; even some of the most distinc- tematic Entomology Laboratory, U.S. National
tive groups within the Eumolpinae are very Museum, NHB 168, Smithsonian Insitution,
heterogenous assemblages (Askevold & Le- Washington, DC 20560 (USNM); Dr. Sharon
Sage 1990). Based on the classification of L. Shute, Department of Entomology, British
Chapuis (1874; still the only one for the Neot- Museum (Natural History), Cromwell Road,
ropical fauna) and which is used by Seeno and London SW7 5BD England (BMNH); Dr.
Wilcox (1982), Glyptosceloides belongs in the Robert L. Davidson, Section of Invertebrate
section Edusites ofthe tribe Eumolpini. Zoology, Carnegie Museum of Natural His-
The characters of Edusites are proepisterna tory, 4400 Forbes Ave., Pittsburgh, PA 15213-
forming ocular lobes, and elytra laterally ru- 4080 (CMNH); Dr. Charles Triplehom,
góse. The only New World members of this Department of Entomology, Ohio State Uni-
section are Argoa Lefévre, Auranius Jacoby, versity, 1735 Neil Ave., Columbus, OH 43210
and Tymnes Chapuis. Argoa, compared with (OHIO); Dr. Shawn M. Clark, División of
Glyptosceloides above, is unlikely to be con- Plant Industry, Dept. of Agriculture, State of
WV
fused with it. Two other New World edusine West Virginia, Charleston, 25305 (SMC).
genera, Auranius and Tymnes, are readily dis- ISAC refers to the collection of I.S. Askevold;
tinguishable from Glyptosceloides. Auranius RWF refers to the collection ofR.W. Flowers.
is monotypic; A. robustus Jacoby is a large
weevil mimic from Brazil. Tymnes is a north- REFERENCES CITED
ern genus, extending south only to Guatemala;
species of this genus lack both toothed pro- Askevold, I.S. in preparation a. An annotated list ofthe
femora and sexual dimorphism of the head. Chrysomelidaeknownto occurinChile (Coleóptera).
Askevold, I.S. in preparation b. A synopsis ofthe genera
Based on comparison with these genera, we
of Eumolpinae and Lamprosomatinae of Chile, with
are confident that Glyptosceloides belongs in comments on higher classification of Eumolpinae and
this assemblage of taxa and that it is also dis- taxonomic revisions of selected genera (Coleóptera:
tinct from them. Chrysomelidae).
Askevold, I.S. & L. LeSage. 1990. A taxonomic revisión
of the genus Jansonius Baly 1878: taxonomic confu-
ACKNOWLEDGEMENTS
sión and tribal reclassification (Coleóptera:
Chrysomelidae: Eumolpinae). Revista Chilena de En-
Research on the part of ISA was supported by tomología 18:29-37.
a Natural Sciences and Engineering Research Chapuis, F. 1874. Histoire naturelle des Insectes: Genera
Council Post-Doctoral Fellowship to ISA and des Coléoptéres. Vol. 10 Famille des Phytophages.
by a grantA(&FLMAX 91005) from CSRS, USDA LiNPDaRrOiTs.H,ivC+.H4.5519p5p7.. The principal terms used for malc'
to Florida University. and female genitalia in Coleóptera. Opuscula Ento-
The foUowing curators or private individu- mológica22:241-256.
áis are owed thanks for loaning specimens in Reíd, C.A.M. 1993. Eboo, nom. nov.: redescription of
type species (Coleóptera: Chrysomelidae: Eumolpi-
the collections under their care, referred in the
nae). Coleopterists Bulletin47(l):61-67.
text above by the corresponding coden: Dr. Seeno, T.N. and J.A. Wilcox. 1982. Leaf Beetle Genera
Mario Elgueta, Museo Nacional de Historia (Coleóptera: Chrysomelidae). Entomography 1:1-221.
Natural, Casilla 787, Santiago, Chile (CHIL); Seeno, T.N., G. Scherer, and K.S. Corwin. 1976. Jan
Dr. Otto Merkl, Természettudományi Bechyné. Necrology and bibliography. Entomologis-
Múzeum, ÁUátra, Baross u. 13, H-1088 Buda- che Arbeitenaus demMuseumG. Frey27:1-69