Table Of ContentRHODORA,
Vol. No. 883/884, 234-253, 1993
95, pp.
CHROMOSOME NUMBER
DETERMINATIONS
IN
FAM. COMPOSITAE, TRIBE ASTEREAE.
V.
NORTH AMERICAN TAXA
EASTERN
Zhang and Chunsheng Xiang
John
Semple,
C. Jie
ABSTRACT
Chromosome numbers are reported for 333 individuals of 109 taxa and two
among
interspecific hybrids from genera. The vast majority of the reports are
1 1
Many
and
for Aster sensu lata Solidago, counts confirm previous reports for the
taxa; approximately one third of the counts are reports for a taxon^ or ploidy
first
level of a laxon, in one or more of the 26 states of the eastern United States and
one Canadian province sampled. The cytogeography and cytotaxonomy of the
The
Aster Solidago and complexes
pitosus, striata S. uliginosa are discussed. fol-
=
=
lowing first reports are included: Aster attenuatus. In 48; A. surculosus, 2n
= = = =
36; Solidago latissimifolia, 2n 2n 36, 2n 2n 54; S. patula var.
18n,
27,i,
= =
strictula, 2n S. pulchra, 2n 18„.
9,^;
Key
Words: Compositae, Astereae, chromosome numbers. Boltonia, Co-
Aster,
nyza, Croptilon, Chrysopsis, Euthamia, Gutierrezia, Heterotheca,
Pityopsis, Solidago, Virgulus, cytogeography, Canada, United States
INTRODUCTION
This
is the fifth in a continuing series of reports by our labo-
ratory and collaborators on chromosome numbers the tribe
in
Astereae (Semple, 1985; Semple and Chmielewski, 1987; Semple,
Chmielew^ski and Lane, 1989; Semple, Chmielewski and Xiang,
Determining
1992). the distribution patterns of cytotypes requires
numerous
counts from the range of a taxon. The counts listed in
paper
this are reported such
as further contributions to studies.
In this paper, our focus is on taxa native to the eastern and central
United and
States adjacent Canada.
AND
MATERIALS METHODS
made
Meiotic counts were from mother dissected
pollen
cells
from buds and
1/EtOH
fixed in the field in glacial acetic acid
3:
:
70%
subsequently stored under EtOH. Mitotic
refrigeration in
made
counts were from from wild
root taken transplanted
tip cells
rootstocks or from grown from the
seedlings achenes collected in
Root
wild. tips were pretreated in 0.01% colchicine or saturated
paradichlorobenzene (PDB) 2-3 Modified
for hr.. fixed in either
234
%
^
al—
Semple Chromosomes
235
1993] et
Camoy's EtOH:
Fixative 1/chloroform
(4:3: glacial acetic acid)
:
or Acetic Alcohol Fixative 1/EtOH: and
glacial acetic acid)
(3:
N
HCl
hydrolyzed in for 30 min. at 60°C before squashing.
1
Both
anther sacs containing pollen mother and meristematic
cells
ft
1%
root tips were squashed in acetic orcein, and counts of chro-
mosomes made
were from prepared Permanent
freshly material.
,
made
slides were in most cases as described by Semple, Brammall
Chmielewski
from
^." made
reported as "2/2 Those counts from late prophase (diplo-
me
meiosis
=
chromosomes Many
were counted, 2n of the taxa
e.g.,
18i|.
studied include polyploids and could be indicated by additional
= =
2n 4x However,
notation, 32. since nearly not
e.g., all, if all
of
these cytotypes appear to have undergone diploidization, counts
them
for are reported without indicating ploidy level.
wat
Vouchers counts deposited with duplicates
for are in
all
made
were
distributed to various other herbaria. Identifications
by Semple. In some the voucher specimens did not agree
cases,
more minor
with published descriptions in one or traits, such as
amount
the of pubescence or color; these cases are indicated
floret
by
Table
the
qualifier "aff." in
1.
AND
RESULTS DISCUSSION
Chromosome
among
from
and two hybrids
representing 109 taxa interspecific
and voucher data
genera are reported in Table All location
1.
1 1
were sampled
Populations
are published here for the time.
first
United
in one province Canada and in 26 states in the States.
in
and goldenrods
In total, 308 of these reports are for asters (Aster,
Two-thirds of the counts con-
and Euthamia).
Virgulus, Solidago
and most presented without
are
firm previous reports for the taxa
comment. Included Table are the following first reports for
in
1
2^-48
from
population)
(one
Lindl,
these Aster attenuatus
taxa:
=
from North
2n 36 (one population)
Texas; A, surculosus Michx.,
population
(one eac
Solidago Mill,
rolina; latissimifolia
=
from Massachusetts
18„ from South Carolina, In 36
= Mas
Rhode In 54
from and Island, fron-
Massachusetts
„
Rhodora
236 [Vol. 95
Table Chromosome number determinations of Astereae from Canada and
1.
= =
Ah Ahmed; Ch G. Chmie-
the United States, arranged alphabetically. Tufail J.
= = Xg =
Su Bambang Agus ChunSheng Xiang.
lewski; 5" C. Semple; Suripto;
J.
When voucher specimens did not a published taxon description in one or more
fit
minor the qualifier "aff." used. State highway numbers are indicated by
traits, is
the standard postal code abbreviation for the state followed by the number. First
'
report for the taxon; - first report(s) for the taxon or ploidy level for the particular
province or
state.
= km
E
Aster acuminatus Michx. In Maine. York Co.: Lyman, ME-1 11 0.1
9ii.
5
ofME-35, 5m
95S7.
c6
& 2n=
US-
A. anomalus Englemann in Torr. Gray. 16. Arkansas.^ Carrol Co.:
W
km S Union
62 of E of AR-23, Su 9950. Co.:
Berryville, 6.9 Illinois.
iSc
&
Bald Knob Rd., woods, S Su 9877,
^
A. asteroides MacMillan (synonym: A. patemiis Cronq.) 2n 18. North
(L.)
W W
km S &
Carolina. Cherokee Co.: of Ranger, US-64 7.8 of US- 129, Jeff
Semple 10353
(2 plants).
=
US-
A. attenuatus Lindl. in Hook.' In 48. Texas. Jasper Co.: E of Woodville,
5*Sm
13.5kmEofUS-69,
190 10055.
2n=
A. avitus Alex. 18. Georgia. Dekalb Co.: base of Stone Mt. near access rd
S &
to top of mountain, Jeff Semple 10346.
&
=
Peawanuck
A, borealis (Torr. Gray) Prov. In 64. Ontario. Kenora Dist.: (S
of old Winisk), Clements
s.n.
=
A. chlorolepis Burgess in Small. In 36. North Carolina. Yancey Co.: Mt.
midway S & Su
Mitchell, along rd to park, 9684.
^
Win-
A. ciliolatus Lindl. 2« 48. Ontario. Kenora Dist.: Peawanuck (S of old
Clements
isk), s.n,
= S & Su
In Union
A. cordifolius L. 16. Illinois. Co.: Trail of Tears State For,,
S & Ch
9882. Kentucky. Allen Co.: US-31 a few mi. S of Petroleum, 9110.
NW
= 5
Pkwy
In 32. Kentucky. Adair Co.: of Toria, Cumberland (MP38.1),
& km
Su KY.1392 Rock-
9447. Pulaski Co.: ca. 10 S of Livingston, bluff of
S&Su
N
castle R., 9603. Oklahoma,- Flore Co.: US-59 just of Ouachita
boundary, S Su
Nat'l For. 9985,
iSc
=
A. aff. cordifolius L. x A. pilosus Willd. 2n 32. Ontario. Waterloo Co.:
Twp, S & Ch
Wellesley 9097
A.
NE
=
of
A. depauperatus (Porter) Fern. 2n 16. North Carolina. Granville Co.:
Butner, along power line right-of-way, S & Su 9724.
NW
= km
A. divaricatus L. 2n Pennsylvania. Perry Co.: PA-74 3 of Ickes-
9,|.
S&Su = N
berg, 9490. 2n Massachusetts. of Somerset along
18. Bristol Co.:
Segreganset Estuary, 5" 9183.
N
2n^
drummondii
A, Lindl. 16. Arkansas. Carrol Co.: edge of Berry ville,
S&Su
AR-21, comer of Julie and Standley Ave. NE, 9948. Louisiana.^
St.
km
Caddo Parish: LA-2 8.0 E of Vivian (LA-1), S Su 10031. Texas.^ La-
iS:
NE
mar S Newton
Co.: Brackeen Cemetery Rd. TX-19, 5w 10003. Co.;
off
c&
W
km =
TX-63 S &
ofBurkeville, 2.5 oTS^hine R., Su 10049. In 32. Illi-
NW
S
Kakankee Rock
nois. Co.: of Bradley, Kakankee State Park, along R.,
&
S &
Su 9867. St. Clair Co.; Fairview Heights, community park, Su 9406.
S
Adams
Mississippi.- Co.: SE of Naiches, Fire Tower Rd. just E of US-91,
4
Semple Chromosomes
al.— 237
1993] et
Table Continued.
1.
& Su 10087. Texas.^- Anderson Co.: US-84 17.6 km E of Palestine city limits
S &
(Loope Rd-56), Su 10016.
=
dumosus
A. L. 2« 8„. North Carolina. Bertie Co.: SE of Bertie, NC-308 9.8
WSW
km
SE of US-17, 5 Su 9742. Yadkin Co.: of Yadkinville, US-421
<$
km S & & S & =
ca. E of US-21, Su 9708 Su 9709. In 16. Kentucky.
1
KY-1392 km S &
Pulaski Co.: ca. 10 S of Livingston, bluff of Rockcastle R.,
W W
km
Su
9604. Louisiana. Concordia Parish, of Ferriday, US-84 6 of LA-
S & N
15, vicinity of picnic area, Su 10069. Massachusetts. Essex Co.: of
W
km S &
Newbury, by airport, ca. 1.5 of Plum R., Su 9574. Mississippi.
Is.
NE
Amite Co.: of Liberty, Holmes Rd. 6.7 km SE of MS-570, S & Su 10102.
= N
In 32. Arkansas. Franklin Co.: of Ozark, AR-23 just S of 1-40, S & Su
W
AR-28 km E km S &
9966. Sebastian Co.: 4.6 of Cauthron, 14 of US-71,
N
km
Su 9980. Louisiana. Vernon Parish: US-171 3.4 of Anacoco (LA-1
1 1),
W
S&Su
km
10041. Oklahoma.2 McCurtain US-70 E of of
Co.; 3.5 Valiant,
& km
Millerton, S Su 9995. South Carolina. Clarendon Co.: US-521 SE
1.1
S&Su
NE
of Wilson, 9784. Texas.= Newton Co.: of Burkeville, TX-63 2.5
W
km &
S Su
of Sabine 10048.
R.,
N
= New
A. gracilis Nutt. 2/2 9„. Jersey. Burlington Co.: of Chatsworth, Co.
km
S
Rd.-563 0.9 S of NJ-72, 10365; S of Chalworth, local road to cranber-
W
W
km
S
ry bog, 10369. Ocean Co.: of Bamegat, Co. Rd.-554 2.4 of Gar-
S &
Su
den State Parkway, 9516.
=
US-98
A. hemisphaericus Alex, in Small. 2rt 18. Mississippi. Franklin Co.:
S&Su
E
of MS-570, 10101.
just
A. laevis L. var. laevis x A. lanceolatus Willd. ssp. lanceolatus var. lanceolatus.
= S & Ch
In Kent Hwy-3, Port Crewe, 9138.
48. Ontario. Co.:
Semple Chmielewski.
A. lanceolatus Willd. lanceolatus var. hirsuticauUs &.
ssp.
= S
2n 32. Wisconsin. Douglas Co.: US-2 E of Maple, 9047-2.
&
Semple Chmie-
(Wieg.)
A. lanceolatus Willd. lanceolatus var. interior
ssp.
= SW AR-23 km E
lewski. 2n 48. Arkansas.' Logan Co.: of Booneville, 2.2
S&Su
White S of Norris
of Carolan Cemetery, 9976. Illinois. Co.: City,
S&Su =
2n Webster
junction of and US-45, 9423. 64. Kentucky.-' Co.;
IL-1
W
S&Su
KY-109 km
of Clay, 9428.
1.7
&
Semple Chmielewski. In
A. lanceolatus Willd. lanceolatus var. latifolius
ssp.
W
S&Su
=
km
49 of Zeiglcr (IL-1
64. Franklin Co.: IL-1 1.7 48).
Illinois.
W
km S
9875. Indiana. Putnam Co.: SE of Cloverdale, IN-42 4.1 of US-231,
W
km
& Su 9401. Kentucky.2 Robertson Co.: KY-617 0.5 of Piqua, banks of
W
S&Su
S&Su km
KY-109 of Clay,
Bluelick 9593. Webster Co.; 1.7
R.,
9427.
&
Semple Chmielewski.
A. lanceolatus Willd. lanceolatus var. latifolius
ssp. aff.
km
In = Montgomery SE of Montgomery, US-321 0.7
Alabama.' Co.:
64.
S&
SE of US-82 Su 10155. The rays are blue and the phyllaries are less
graduated than typical for the variety).
km
= US-60 E of Van
A. 2n Missouri, Carter Co.: 1.8
laterijlorus (L.) Britt. 16.
N
S&Su
US-59 of Ouachita Nat'l
Buren 9907. Oklahoma. Flore Co.: just
= E
& Alabama. Dale of
For. boundary, bank, 5 Su 9984. 2« 32. Co.:
river
S&Su
10152.
US-84 Choctawhatchee R., Illinois.
Clayhatchce,
at Little
Rhodora
238 95
[Vol.
Table Continued.
1.
km S d
Hamilton IL-142 SE of Dahlgreen, Su 9419. Kentucky.
Co.; Estill
1
N
km
Co.: KY-52 E of Ravenna, S & Su 9457. Hopkins Co.: KY-i09 of
1
km
Charleston, S & Su 9430. Pulaski Co.: KY-1392 ca. 10 S of Livingston,
MO-
bluff slope of Rockcastle R., S & Su 9601 Missouri. Douglas Co.: 14 at
S &
Twin Bridges (MO- 18 along N. Fork of White R., Su 9927. Tennes-
1),
km &
see, Montgomery Co.: US-79 0.5 S of Kentucky state line, S Su 9435.
Texas. Rusk Co.: US-84 6.4 km E of Reklaw, S & Su 10023.
N
=
A. linariifolius L. In Louisiana. Rapides Parish: of Forest Hill, LA-1 12
Qn.
W NW
km & Popham
2.1 of US-165, S Su 10090. Maine. Sagadahoc Co.: of
Beach, junction of ME-209 and ME-216, S 10378. Massachusetts. Barnstable
km NE &
Co.: Harwich, Queen Anne Rd 2 of Old Chatham Rd., S Su 9559.
W
New Jersey.2 Atlantic Co.: US-40 5.6 km of Mays Landing (NJ-42), 5 &
km
Su 9504. South Carolina. Berkeley Co.: SC-45 5 E of Stephen,
ca. St.
S &
Marion Su
Francis Nat'l For., 9794.
W
=
Popham ME-209
A. nemoralis Ait. In 9„. Maine. Sagadahoc Co.: Beach,
2kmEofME-216,
S70i50.
=
A. novi-belgii L. In 48. Maine. York Co.: Wells, vicinity of Municipal Of-
5
5w
9581.
fices, c6
=
A. ontarionis Wieg. In 32. Tennessee. Marshall Co.: Henry Horion State
S&Ch
NE km
Montgomery
Park, 9117. Co.: of Clarkville, US-79 0.5 S of
S &
Kentucky Su
state line, 9436.
=
A. oolentangiensis Riddell. 2n 32. Illinois. St. Clair Co.; Fairview Heights,
community S & Su MO- km
park, 9407. Missouri. Oregon Co.: 19 16.6 S of
N S&Su
km
US-60, of Greer, 9911. Wayne Co.: Co. Rd.-F 0.1 E of US-67,
km
&
S
7.8 S of Greenville, Su 9894. Oklahoma. McCurtain US-259
Co.:
km N S
20.9 of Broken Bow, Su 9990,
iSi
&
A. aff. parviceps (Burgess) Mackenzie Bush. [See text for discussion of these
W
=
MO-8 km
collections.] In 32. Missouri. Washington Co.: 6.6 of Potosi,
Su & Ah
9393.
S.
A. aff. pilosus Willd. var. pringlei (A. Gray) Blake. [See text for discussion of
- N MO-
these collections.] 2n 48. Missouri.' Oregon Co.: of Greer, 19 16.6
km
S of US-60, Mark Twain S & Su
Nat'l For., vicinity of For. Rd.-3174,
W
9912. Taney Co.: MO-76 12.6 km of county cedar S & Su
line, glade,
Wayne km km S
9938. Co.: Co. Rd.-F 0,1 E of US-67, 7.8 S of Greenville,
d
Su
9892.
=
A. pilosus Willd. var. pilosus. 2rt 32. Ontario. Essex Co.: Colchester N. Twp,
SW
km SW
Essex-12 S &
5 of Gesto, of Canard Ch 9145. Arkansas-
R.,
Madison Co.: S of Eureka AR.23 km S & Su
Spring, S of county
1.8 line,
km
9955. Illinois. 7.5 E of McLeansboro Co. Rd.-1400E, S & Su 9422.
at
Kentucky. Nicholas S & Warren
Co.: Bluelick Su 9596.
Battlefield State Park,
US-68 Logan &
Co.: at Co. line, vicinity of quarry, S Su 9441. North Caroli-
Avery E S &
na. Co.: of Linville, top of Grandfather Mt., edge of parking lot,
=
Su In km S
9672. 48. Illinois. Hamilton Co.: IL-142 1.0 SE of Dahlgreen,
W
&
Su km
9417. Oklahoma.^ McCurtain E of
Co.: of Millerton, US-70 3.5
S
Su
Valiant, 9994.
iSc
=
A. pilosus Willd.
aff. var. pringlei (A. Gray) Blake. In 48. Massachusetts.
W
Barnstable Co.: East Brewster, along Bike Trail of Nickerson State Park,
i
Semple Chromosomes
al.—
1*^93] et 239
Table Continued.
1.
near MA-6A, S & Su 955 L (The more
plant is hairy than typical but not as
hairy as var. pilosus.)
= NE
* A. praealtus Poir. var. praealtus. In 32. Mississippi. Amite Co.: of Liber-
S&Su
Holms
Rd., 10105. Missouri. Greer SE of Walnut Grove,
ty, Co.:
MO-
Farm S &
123 Rd-56, Su 9946. Oklahoma. McCurtain US-70 E
at Co.:
^
edge of Idabel, S & Su 9993. In = 48. Texas, Lamar Co.: US-27 S of Pow-
1
N
N
dcrly, of Paris, of Loop-Rd.-287, 5<fc 5w 9999.
^
* A. prenanthoides Muhl. In 32. Kentucky. Laurel Co.: along Rockcastle R.
S &
above mouth of Blair Branch Creek, Su 9854. Menifee Co.: Boone Nat'l
S &
KY-715 Su E
For., along river, 9465. Tennessee.- Unicoi Co.: of Unicoi,
t
km
TN-107 5.2 E of TN-173, 5 & Su 9632.
= N
A, priceae Britt. In 64. Kentucky.- Logan Co.: Russellville, LIS-68 just of
* S &
US-431, glade, Su 9438. Warren Co.: US-68 Logan County by
at line,
c\udiVvy,S&Su 9440-1.
W
2n= km
Rabun US-76 S
A. punicens L. 16. Georgia.- Co.: ca. 3 of Cia>ton,
,
& Su MN-18 N
9830. Minnesota. Pine E of end of Big Pine Lake, near
Co.;
W
county 5 North Avery of Grandfather
9069. Carolina. Co.: Linville,
line,
Mt., near Black Rock Hiking Trail, S & Su 9673. Tennessee.- Unicoi Co.: E
W
NC
TN-107 km S & Su
of Unicoi, 0.3 of state line, 9642.
name
A. racemosus [Specimens treated in most floras under the misapplied
Ell.
*
= N
Chowan NC-37
A. vimineus Lam.] In North Carolina. Co.: jusi of
8„.
=
Albcrmarlc Sound Causeway, S & Su 9747. In Alabama.- Montgomery
16.
^
km &
Co.: SE of Montgomery, US-231 0.7 SE of US-82, S Su 10156. Tusca-
W W NW
km S
loosa Co.: US-82 2.4 of US-43, of Northport, of Tuscaloosa,
N
& S
Su 10162. Arkansas. Washington Co.: US-71, rest area of Brentwood,
'
W
& km S &
Su 9959. Kentucky.^ Webster Co.: KY-109 1.7 of Clay, Su
km
LA-67 LA-
9429. Louisiana.- East Feliciana Parish: S of Felps, 2.5 S of
,
N
km
LA-7
422. Su 10107. Red River Parish: of Edgefield, 1.2 S of
.V rf 1
NW
county S & Su 10036. Mississippi. Franklin Co.: of Roxie, US-84
line,
W
km
km of MS-33, S & Su 10098. Monroe Co.: S of Becker, MS-25 0.9
1.5
N km
of MS-8, S & Su 10167. Missouri.^ Wayne Co.: Co. Rd.-F 0. E of
1
km S d Su Anderson US-84
US-67, S of Greenville, 9895. Texas. Co.:
7.8
km S & Su Grimes
17.6 E of Palestine city limit (Loop Rd-56), 10017. Co.:
W
km & TX-150
TX-105 2.6 of Plantersville, Su 10064. San Jacinto Co.:
.S"
3.6 km E of Shepherd, S & Su 10062. 2n - 32. North Carolina. Durham
Co.: N of Durham, by Eno R., Penny's Head Nature Res., S & Su 9718.
=
A, rctiailatus Pursh. 2n 18. Florida. Leon Co.: S of Bloxham, Appalachicola
R4W, S & Sem-
N.F., For. Rd.-320 just E of Co. Rd.-375 (TIS, Sec. 32), Jeff
ple 10342.
= NC-226
DC. In North Carolina. Mitchell Co.: 0.5
A. 48.
retroflexus Lindl. in
W
km E of Tennessee S & Su 9648. Watauga Co.: of Blowing
state line,
km 5
Rock, S &Su 9681. Tennessee. Unicoi Co.: TN-107 17 E of Unicoi,
<fe
Su
9643.
=
Schum. In South Carolina.^ Chesterfield
A. sericocurpoides (Small) K. 18.
NE S & Su
Co.: US- of entrance Cheraw Fish Hatchery, 9780.
just to
1
NW
-
Kakankee of Bradley,
A. Hook. 2n Co.:
shortii Lindl. in 16. Illinois.
S &
Kakankee woods by Rock Su 9868. Kentucky. Adair
R. State Park, R.,
Rhodora
240 [Vol. 95
Table Continued.
1.
W
NW
S & Su
Cumberland Parkway (MP38.1) of KY-80,
Co.: of Toria,
W
= km
9449. In 32. Indiana. Putnam Co.: SE of Cloverdale, IN-42 4.1 of
S&Su
KY-109 SE Dawson
US-231, 9402. Kentucky. Christian Co.: of
Spring, Pennyrite For. State Park, S & Su 9433. Warren Co.: S of Alvaton,
km
5
US-231
2.8 S of Co. Rd.-961, C/z 9105,
c£
km
2n=
A. solidagineus Michx. 18. North Carolina. Cherokee Co.: US-64 7.8
W
5
of Ranger (US- JeffSemple 10352
129), <& (3 plants).
W
=
A, spectabilis Ait. In 36„. Massachusetts. Plymouth Co.: of Cedarville,
N N
km S
Long Pond Rd. 0.3 of Carters Bridge Rd., of Great Herring Lake,
&
Su
9567.
= New
2n Ocean Tuckerton,
A, subulatusl^. var. subulatus. 5„. Jersey. Co.;
&
cedar marsh E of town center, S Su 9525.
=
A, surculosus Michx.' In 36. North Carolina,- Cherokee Co.: SE of Ranger,
W
km
S &
US-19
2.35 S of US-64, of Ranger, JeffSemple 10350.
W
= km
A. texanus Burgess in Small. In 32. Texas. Grimes Co.: TX-105 2.6
S &
Su
of Plantersville, 10065.
=
Farm
A, texanus Burgess in Small. 2n 32. Texas, Jasper Co.: US- 190 at
afT.
W
S d
Rd-256, of TX-63, Su 10054.
- km
Michx. 2n South SC-652 SE
A, tortifolius Carolina. Jasper Co.: 14.9
9,|.
S &
Su
of Pineland, 9804.
W
= km
MO-8
A. turbinellus Lindl. 2n 96. Missouri. Washington Co.: 6.6
ca.
&
of Su Ah
Potosi. 9392.
5,
=
New
A. umbellatus Mill. var. umbellatus. 2n Connecticut.- London Co.:
9i,.
Waterford, Waterford Conservation Area, Ncvins Brook, S 9191. Massachu-
W
N
km
setts.2 Essex Co.: of Newbury, ca. 1.5 of Plum R., opposite air-
Is.
S&Su
port, 9573.
= km
A, undulatus L. 2n 9„. South Carolina, Berkeley Co.: SC-45 ca. 5 E of St.
=
Stephen, Francis Marion Nat'l For,, S & Su 9793. 2n 32. Kentucky. Adair
NW
W
&
Co.: of Toria, Cumberland Parkway (MP38.1) of KY-80, S Su
9448. North Carolina. Watauga Co.: US-421 at Wilkes Co. line, S & Su
9701.
DC.
A. urophyllus Lindl. 2n
in Berryville
km
US-62 6.9 E of AR-23, S d Su 9949. Minnesota. Hennepin Eden
Co.,
W
Prairie, Spring Rd. just of county airport, S 885 L Missouri. Carter
et
al.
SW
Co.: between Hunter and Van Buren, gravel extension of MO-T 7.8 km
S&Su
km
of US-60, 9903. Hickory Co.: E of Wheatland, US-54 0.9 E of
Hicomo S & Su Wayne MO-49
Picnic Area, 9943. Co.: E of Williamsville,
km S&Su
E
4.4 of Co. Rd-A,
9901.
=
&
Boltonia astewides (L.) L'Her. var. recognita (Fern. Griscom) Cronq. 2n
km
18. Illinois. Hamilton Co.: IL-142 SE of Dahlgreen, S & Su 9415\ IL-
1
km S&Su
142 8.5 SE of Dahlgreen,
9420.
Conyza =
canadensis Cronq. van Waterloo Re-
(L.) canadensis. Ontario.
2/7 18.
gional Mun.: Waterioo, University of Waterioo, north campus greenhouse,
S
weed
in pot, 1038x.
W
=
Croptilon divaricatum Raf
(nutt.) 2n North Carolina. Pitt Co.: of
4,,.
km S&Su
US-64 E
Bethel, Edgecombe
0.2 of Co.
9736.
line.
Semple Chromosomes
al.—
1993] et 241
Table Continued.
1.
=
Chrysopsis gossypina (Michx.) Ell. ssp. gossypina gossypina. In 9„. North
f.
W
km
Carolina. Pender Co.: Hampstead, NC-210 0.9 of US- S & Su
17,
9758.
=
C. gossypina (Michx.) Ell. ssp. gossypina trichophylla (Nutt.) Semple. In
f.
km
9„. North Carolina.^ Bladden Co.: NC-53 20 SE of US-701, S & Su
9773.
W
mariana = New km
C. 2n US-40
(L.) Ell. I2„. Jersey. Atlantic Co.: 5.6 of
Mays S &
Landing (Hwy-52), Su
9503. Louisiana. Beauregard LA-i:
Parish:
W W
km
km
of Ragley (US- of Gordon Tower, S & Su
1.8 171), 1.4 Fire
1
km
10089. North Carolina. Bertie Co.: SE of Bertie, NC-308 9.8 SE of US-
NW
\l,S&Su
9742. South Carolina. Colleton Co.: of Jacksonboro, NC-64
NW
km 5 &
US- Su
ca. 2.5 of 9799.
17,
& =
Euthamia
leptocephala (Torr. Gray) Greene. 2« 18. Texas. Navarro Co.:
W
W
km &
of Eureka, US-287 3.3 of Farm Rd.-637, S Su 10010. Newton
W
TX-63 km NE S &
Co.: 2.5 of Sabine R., of Burkeville, Su 10050. Polk
Soda
Su
10061.
=
NC-56
E. tenuifolia (Pursh) Nutt. 2rt 18. North Carolina. Franklin Co.: 4.2
W
km
S & Hampton
of county Su 9731. South Carolina.^ Co.: SC-68
line,
NW
km &
of Yemasee, S Su 9802.
1 1
& =
Gutierrezia texana (DC.) Torr. Gray var. texana. 2n 4„. Texas. Dallas Co.:
N
RR
S &
of Cedar Hill, Cedar Hill Rd. opposite lot No. 10, right-of-way,
1 1
Su
10008.
=
camporum
Heterotheca (Greene) Shinners var. glandulissimum Semple. In
N
S &
36. Alabama.- Limestone Co.: Littleville, US-43 at city limits, Su
NE
Oak AR-21 S
10178. Arkansas. Carrol Co.: of Grove, just S of Blue Eye,
& Su Ozark, MO-14 E of US-65, S & Su
9947. Missouri.2 Christian Co.: just
9939.
W
= New 5
H. Lam. In Salem of Malaga, NJ-55,
subaxillaris Jersey.- Co.:
9i,.
&
Su
9498.
=
Cape
Pityopsis falcata (Pursh) Nutt. In 9„. Massachusetts. Barnstable Co.:
W
MA-6A,
Cod, East Brewster, by Bike Trail of Nickerson State Park, near
S&Su
9762.
& =
Semple Bowers. In
P. graminifolia (Michx.) Nutt. var. latifolia (Fern.) 18„.
N
North Chowan NC-37 of Albermarle Sound Causeway,
Carolina. Co.: just
W
km
SE of Edenton, S & Su 9746. Pender Co.: Hampstead, NC-210 ca. 4.1
5
of US- 5m
9762.
1 7, <S
=
&
Semple Bowers. In
P. graminifolia (Michx.) Nutt. var. tenuifolia (Torr.)
S &
LA- SE of DtQumcy, Su 10083.
9„. Louisiana.' Beauregard Parish: 12
km S
East Feliciana Parish: E of Woodland, LA-432 1.5 E of LA-67, Su
<&.
10113.
W
=
P. oligantha (Chapm.) Small. In 36. Florida. Franklin Co.: of St. Teresa,
US-319 2.7 km S of Ochlockonee R., 5 & Jeff Semple 10340.
= TX-105
Grimes
Solidago altissima L. var. altissima. In 36. Texas.- Co.: 1.5
N
km
E of TX-Farm Rd-362, S & Su 10069. Harris Co.: of Bayton, I- 10 rest
Rhodora
242 95
[Vol.
Table Continued.
I.
N
S & Su 10076. Lamar S of Powderly, US-271 of Loop Rd-286,
area, Co.:
S&Su
= km
10002. 2n 54. Illinois. Hamilton Co.: IL-142 SE of Dahl-
1
S & Su KY-109 SE Dawson
green, Q41S. Kentucky. Christian Co.: of Spring,
N &
S Su US-62
end of Pennyrite Forest State Park, 9434-1. Harrison Co.: 2
SW N
km
of Clay S & Su 9592. Louisiana. Vernon Parish: of Anacoco,
ville,
N NE
km
US- 171 3.4 of LA-1 5 Su 10046. Missouri. Washington Co.:
1 1, <fe
N
of Potosi, .MO-2I just of MO-8, S, Su & Ah 9395. North Carolina. Jones
W
km NE S &
Su
Co.: Chadwick, US-17 ca. 5 of Maysville, 9753. Pitt Co.:
W
km S &
of Bethel, L'S-64 0.2 of county line, Su 9737. South Carolina.'
NW
km &
NC-64 Su
Colleton Co.: of Jatksonboro, 2.5 from US-17, S.
km
9800. Tennessee.- Montgomery Co.: US-79 0.5 S of Kentucky state line,
NE
km &
S Su
ca. 2 of Clarkville, 9437.
1
NW
= km
S. arguta Ait. ssp. arguta. 2w Pennsylvania. Juniata Co.: 5 of
9,,.
S &
Su
Ickcsbcrg. ridge top, 9485.
=
S. arguta Ait. ssp. caroliniana (A. Gray) G. Morton. In South Carolina.
9,,.
km
Berkeley Co.: SC-45 ca. 5 E of St. Stephen, Francis Marion Nat'l For., 5"
& = NE
Su 9792. 2« 18. Mississippi.' Amite Co.: of Liberty, Holmes Rd.
6.7 km SE of MS-570, S & Su 10104. Missouri.' Taney Co.: MO-76 1.8 km
1
W
NW
S & =
of county Su 9935. In Kentucky. Adair of Toria,
line, 36. Co.:
&
Cumberland Parkway S Su Boone
{MP38.1), 9444. Laurel Co.: Daniel St.
For. junction of For. Rd.-l 19 and For. Rd-56, S & Su 9613.
=
S. bicolor L. 2n 9„. Maine. Sagadahoc Co.: S of Bath, Meadowbrook Rd.,
Perry
NW
km
S &
74 of Su
3 Ickesberg, 9487.
W
= km
cacsia L. 2a7 IS. Indiana. Putnam Co.: SE of Cloverdale, IN-42 4.1
.*?.
S &
of US-:31. Su 9403. Kentucky, Powell Co.: Boone Nat'l For. Tunnel
N
Ridge Rd.. .V Su 9462. North Carolina.' Durham Co.: of Durham, Pen-
<.^
ny's Bend Nature Reserve, Eno R., S & Su 9720.
=
canadensis In Plymouth E of
S. L. var. canadensis. Massachusetts. Co.:
9^j.
MA-
Raynham.
S
104 just S of Lake Nippennicket, 9181.
W
= km
S. delicaiula Small. 2« Texas. Anderson Rd.-494 of
IS. Co.: Co. 0.5
W
km
&
S-287, of Cayuga, S Su
I 3.5 10014.
W
= New km Mays
S. crecta Pursh. US-40 of
2/i 9„. Jersey. Atlantic Co.: 5.6
W
& Su km
Landing, 9501. .North Carolina. Franklin Co.: NC-56 4.2 of
.S'
S & = KY-
county line, Su 9735. 2n Kentucky.^ E of Ravenna,
18. Co.:
Estill
km
52 by Ravenna- Pryse Rd., S d Su 9454. Pulaski Co.: KY-1392 ca. 10 S
of Livingston, bluff of Rockcastle R., S & Su 9605. Mississippi.' Itawamba
N
MS-23 km S &
Co.: ofTremont, Su
10.1 10175.
W
=
S.,tistulosa Mill. 2a? 9„. North Carolina. Martin Co.: US-64 just of Par-
S&Su
N
=
melc, 9739. In Alabama.' Mobile of Theodore, Co. Rd-
18. Co.:
km N
59 ca. 2 S of I-IO, S & Su 10128. Mississippi.' Harrison Co.: of
N
MS-67 S New
D'lberville, of 1-10, 5« 10125. Jersey. Atlantic Co.:
c6
swampy
woods S &
b> NJ-50 just S of US-40, Su 9506. North Carolina.
Jones Co.: Chadwick, US-17 ca. 5 km NE of Maysville, S & Su 9751. South
Carolina. Clarendon US-521 km SE 5 5m
Co.: of Wilson, 9785.
i.l
cS
=
S. JJe.xicauUs L. 2n 36. Kentucky. Estill Co.: KY-52 E of Ravenna on Raven-
S &
na-Pr}se Su
Rd., 9453. Menifee Co.: Boone, N.F., KY-715, river bank.
.
al—
Semple Chromosomes
1993] et 243
Table Continued.
1.
S&Su
KY-1392 km
9464. Pulaski Co.: ca. 10 S of Livingston, bluff of
S d
Rockcastle Su
9606.
R.,
=
k 2n
S. aff./lexicauHs L. 90.' Kentucky. Leslie Co.: S of Stinnett, US-421 just S
of where highway S & Su
crosses river, 9619. [The plant was for the spe-
tall
and
cies the heads were very
large.]
=
S. gigantea Ait. In 18. Kentucky,^ Leslie Co.: US-421 S of Stinnett, S & Su
N
9620. Mississippi. Lowndes Co.: Columbus, end of tow, US-45 S of Chap-
W
man S & Su km
Rd., 10165. North Carolina. Franklin Co.: NC-56 4.2 of
Nash County S & Su 9732. Washington E of US-64
line, Co.: Skinnerville,
W
S&Su
N
just of NC-32, 9748. South Carolina.- McCormick Co.: 4 km of
S&Su
=
McCormick,
along side rd off SC-28, 9819. In 36. Illinois. Kakan-
NW
S&Su
kee Co.: of Bradley, Kakankee R. State Park, 9871. White Co.: S
S&Su
of Norris City, junction of IL-1 and US-45, 9424. Missouri.^ Douglas
S&Su
MO-14
Co.; Twin Bridges (MO-181), woods by N. Fork White
al R.,
km
9928. Hickory E of Wheatland, US-54 E of Hicomo
Co.: ca. 0.9 Picnic
S&Su N
=
Area, 9945. In 54. Nebraska.^ Hooker Co.: of Mullen, NE-97
at
&
Middle Loup Su Ah
R., 9200.
S,
W
=
glomerata Michx. 2n North Avery of
S. 108. Carolina, Co.: Linville,
SW
&
S
Grandfather Mt., cliff face at top of mt. of swing bridge parking lot,
S&Su
Su 9670; 5000 9676, Mitchell Co.: top of Roan Ml., Roan
ca.
ft. el.,
S&Su
High 6250 Yancey
Bluff, el, 9661. Co.: Mt. Mitchell, near top, ca.
ft.
S&Su S&Su
6000 midway mid
9685; along rd. lo park, 9683.
ft. el., el.,
NW
& -
S. gracillima Torr. Gray. 2n 9„. South Carolina.- Barnwell Co.: of
N
S &
US-278 Su
Barnwell, just of Salkehatchie R., 9814.
=
S. hispida Muhl. 2/2 18. Ontario. Kcnora Dist.: Peawanuck, Clements s.n.
S.jtincea Ail. In ^ 18. Illinois.^ Jefferson Co.: S of Mt. Vernon, 1-64, S & Su
km S &
9414. Effingham Co.; Shelbyville, 2.5 S of Sigel, Su 9H74. Massachu-
5
setts. Bristol Co.: vicinity of Horseneck Beach Park, 9187. Plymouth Co.:
E of Raynham, MA-104 S of Lake Nippeniokct, S 9182.
just
& ^ N
S. houghtonii Torr. Gray. 2n Ontario. Bruce Co.: Bruce Peninsula.
9,1.
S
of Dyer Bay, alvar below Cabot Head, 10363,
^
S. latissimifoUa Mill. 2n ca. 18^,.' South Carolina. Chesterfield Co.: US-1 just
SW
&Su =
S Plymouth
of US-52, 9778. 2n 36.' Massachusetts. Co.: Center
=
N
HillRd. km of MA-3 S & BrouH/et 3553. 2n 27„.' Rhode Island.
2.6 A,
W
W
Washington E of Charleston, of Matunuck, near end of Cards
Co.:
W
S&
=
Pond Plymouth Man-
Rd., Su 9544. 2n 54.' Massachusetts. Co.: of
S&Su
Dam km MA-i
omet, Beaver Rd. 0.9 S of A, 9568,
& =
S. leavenworthii Torr. Gray. 2n 36. South Carolina. Allendale Co.: Yen-
S&Su
ome, US-278
S of Barnwell, 9807,
=
S. lepida DC. Ontario.- Kenora Dist: Peawanuck (S of old Winisk),
2/7 18.
Clements Clements
s.n.-l, s.n,- 2.
km
= Bend TX-36
S. ludoxiciana (A. Gray) Small. 2w 18. Texas.^ Fort Co.: 5 S
of county Hne, N of Farm Rd.-1489, S & Su 10074. 2n = 36. Texas.
just
W
km 5 & Su
Grimes Co.: TX-105 2.6 of Plantersville, 10066. Polk Co.:
5^
US- 190 E of Livingstone, 1.8 km E of Soda (16.4 km E of TX- 146), 5m
70055.
NW
= Hum-
5^. missouriensis Null. 2n South Dakota. Minnehaha Co.: of
9,,.
