Apterous Lathrobium (Coleoptera, Staphylinidae) from the Kii Peninsula in Japan : 3. Group of Lathrobium brachypterum

2006·9.4 MB·

by  Watanabe, Y

E:/、・f,・a,Tok、・),34(1):47-62,May20、2006
ApterousLath1,obium(Coleoptera,Staphylinidae)from
theKiiPeninsulainJapan
3.Groupof !f/?''o/)ltfn1フf1'acﾈvpfe''t'
Yasuaki WATA1、ABE
LaboratoryofInsectResources,TokyoUniversityo、fAgriculture、Atsugi,Kanagawa243-0034Japan
Abstract SevenapterousspeciesofthestaphylinidgcnusLat/1''obiit1'1fromthe
KiiPeninsulaincentralHonshu.Japanarcdealt、、,ith.Allthespeciesbelongtothe9「cuP
ofL(s.sir)bl-ac/1、l)te1-t1l11;sixofthemarcdescribedasno、vtoscienceunderthenamesL.
(ssir)ko、・asal11l111.L.(s.str)o/1to/1e11.・le.し.(s.str)1se11se,L.(s.str)nankiense,L.(s.st「.)
k111o/u川elseandL.(s.str)、、,al,a111ata川tm,andtheremainingonedeterminedasL.(s.st「.)
l111t.,;tielisrecordedwithcollectingdata.
ThemembersofthegroupofLat111o/)111mb1-(1ch、pte1・urnaresimilarinfaciesand
colourtothoseofthegroupsofL.shingo11andL.po11e11s.Theyare,however,distin-
guishablefromthefattersbytherelativelysmallerbodyandsecondarysexualcharac-
tersofabdominalsternitesinthemale,the8thsterniteofwhichistruncateorslightly
emarginateatthemiddleofposteriormargin.
AsthememberofthegroupofLbr(1chypte1-uni,onlyonespecies.L masao1,has
hithertobeenreported fromthe Kii Peninsula(WATANABE,1999,p ie9). Fo11oWin9
thetwopreviouspapers(WATANABE,2005a,b),IamgoingtodescribethenewSpecies
belonging to thegroupof L bl・aclypte1・uln inaddition to thecollectingdataof L
masao1.
Beforegoingfurther,IwishtoexpressmyheartythankstoDr.Shun-lobiUENo,
visitingProfessoratTokyoUniversityofAgriculture, forhiskindnessingivingme
valuableadviceonthepresentstudy.DeepgratitudeisalsoduetoDr.ToshioKISHI-
MoTo,JapanWjldlifeResearchCenter,Messrs. IsaoMATOBA,WakayamaPrefeCtu「al
Museum ofNatural History, Hiroki SAT0, Matsuidamachi, Gunma, and Hideyuki
YoKozEKl,Yokkaichi-shi,Mie, fortheirkindnessinsupplyingmewiththespecimens
usedinthisstudy,andMessrs.ArataIsHlzUKA.JapanWildlifeResearchCenter.Junno-
sukeKANToH, LaboratoryofInsectResources,TokyoUniversityofAgriCultu「e, fo「
theirassistanceintakingthephotographsinsertedinthispaper.
Lat/1robium(s.str.)masaoiY.WATANABE,1999
[Japanesename:Kii-himckobanc-nagahanckakushi]
Lath/・obilm1(s.str),11asao1Y.W、T、NABE,l999.Ent.Rev.Japan、Osaka,54:109-112
4S YasuakiW/、T^N^Ill
ThisSpecieswasdescribedbyWATANABE(1999,p.109)basedonsixmaleand
fOu「femalespecimensobtainedonMt.Wasamata-yama inNaraPrefecture. since
then,thisspecieshasnotbeenreporteduntilnowfromotherlocalitiesofJapan.Itjs
newtothefaunaof MiePrefecture.
S eel'nons e_、-emfnee/. 2 2 Hirakura, Mie Pref., Honshfi, Japan,
. ,
24-III-1992,Y.WATANABEleg.
1sfr!0zlflon. Japan(centralHonshu:KiiPeninsula).
Lat/1「obium(S.str.)koyasallumY.WATANABE,spnov
[Japanesename:Koya-himckobane-nagahanckakushi]
(Figs.1、2,8-10)
Bodylength: 6.4-7.6mm(fromfrontmarginofheadtoanalend);3.2_3.4mm
(fromfrontmarginofheadtoelytralapices).
Bodyelongate, nearlyparallel-sidedandsubdepressedabove.Colour reddjsh
browntobrownishblackandmoderatelyshining,withmandiblesandantennaebrown_
ishre palpi,labrum,legsandtwoapicalabdominalsegmentsbrownishyellow.
ThepresentnewspeciesmaybeplacednearLmasao1fromMt.Wasamata_yama
ofNaraPrefectureinviewofgeneralconfigurationofthemalegenitalorgan,but js
distinguishablefromitbythefollowingpoints:
Maleand female. Headsubquadrate,lesselevatedmediallythaninL
masao1',
Slightlytransverse(width/length=1.03)oraslongasbroad,widestatposterjor fourth
andmoredistinctlynarrowedanteriadthaninLmasao1;lateralsidesgentlyarcuate;
Su「faceSlightlymorestronglyandmorecoarselypuncturedthaninLmasao1'though
Cove「edwithsimilarmicroscopicgroundsculpturetothatofL nlasa()1; postocular
Pa「tIon9erthaninLmasao1,about3.14timesaslongaslongitudinaldiameterofeye.
Antennaeelongate,extendingalittlebeyondthemiddleofpronotumandnotthjck_
enedtowardstheapicalsegment, twoproximalsegmentspolished, theremajnjngs
opaque, 5thto 10thsegmentsmoreor lessmoniliform,andsimilar jnartjculatjonto
thoseof
,71asao1.
.
P「onOtum similar inconfiguration tothatof L nlast1o1,apparently longer than
b「cad (length/width=1.23), distinctly longer (pronotum/head=1.38) and a ljtlle
b「oader (Pronotum/head=1.10) than head; surface sparingly and slightly more
COa「SelyPuncturedthaninLnlasao1exceptforanarrown、ediansmoothspaceElytra
Subt「aPezoida1,somewhtatmorestronglydilatedposteriadthaninL
masao1, appar_
entlyt「anSverse(width/length=1.19),distinctlyshorter(elytra/pronotum=078)buta
littleb「oader(elytra/pronotum=1.14)thanpronotum;lateralsidesnearlystrajght,pos_
te「iOrmarginbroadlyemarginateatthemiddleasinL masaot;surfacesparjnglyand
me「e「eu9hlypuncturedthaninLmasaot.Legsmoderatelylong,profemora,protjbjae
andprotarsisimilarinstructuretothoseofthemembersofthisspecies_group
Abdomeneton9ate,somewhatdilatedfrom3rdto6thsegmentsandthenabruptly
na「「owed towardsthe anal end as in L masa()1;eachtergitesimilarlypuncturedand
AptcrousLat/1・l()1/11‘In「romtheKiiPeninsula 49
Fjg l Lat1/,・obltl111(s.sir)、/-olasantl111YW、T、、、、、BI:,sp nov.,(S 「romMt.Koya-sanofWakayamaPre-
fecture,Japan.Scale:1.0mm.
pubescentasinLmasao1;inmale,8thsterniteshallowlythoughdistinctlyemarginate
atthemiddleofposteriormarginandsomewhatdepressedbeforetheemargination.
surfaceofthedepressionmorecloselyprovidedwith neblackishbrownsetaethanin
otherparts,71hsterniteaml ostsmi ple,infemale,8thsterniteabruptlyna「「owedin
posteriorthirdstowardsthegentlyroundedapex.
MalegenitalorgansimilaringeneralconfigurationtothatofLmasao1indo「Sal
ll in the followingpoints: ventral scleriteofmedian lobe
vjew, but djfferent from
clubbedandmuchmoreelongate; fusedparameresymmetrical,uniformlyna「「owed
towardstheapex,dorsalsurfaceprovidedwithafinelongitudinalcarinaatthemiddle
ofposteriortwo-thirds.
50 Yasuaki WATANABE
5
Figs.2-7. Secondarysexualcharactersofabdominalsternitesinthemale;L.(s.sir)ko、,asanun1sp nov
(2);L.(s.str)ohto1/ensespnov.(3);L.(s.str)!se11sespnov.(4);L.(s.str)flail、・/fells.osp nov.(5),L
(s.str)kinokunte,1sespnov(6);L.(s.str)、、,astlnlatanu111spnov.(7).
Typeselies. Holotype: ,allotype: Mt.Koya-san,WakayamaPref.,Honshu,
,
Japan,11-XI-i971,Y.WATANABEleg.Paratype:1(3,samedataasfortheholotype.
Thetypespecimensaredepositedin thecollectionof theLaboratoryof InsectRe_
sources,TokyoUniversityofAgriculture.
Distribution. Japan(centralHonshLl:KiiPeninsula).
Rema;'ks. This new species is also similar in general appearance to L.
blaChyPtelumSHARP(1889,p255),butarereadilydistinguishablefromitbyconfjgu_
rationofthemalegenitalorgan.
Bionomics. Allthetypespecimenswerefoundintheleaflitteraccumulatedjna
deciduousbroadleavedforestonMt.Koya-san.
Etymology. Thespecificepithetofthisnewspeciesisderivedfrom''Mt Koya_
san”,thetypelocality.
Antcrous1_at/11・o/111111「romthe KiiPeninsula
Figs.8-10. MalegenitalorganofLa1/11obium(s.str)ko、asantl1nsp nov;dorsalview(8),lateralview
(9),andventralview(10).Scale:l.0mm.
Lathrobium(s.str.)ohto/1e1'seY.WATANABE,sp nov
[Japanesename:etc-himckobane-nagahanekakushi]
(Figs3,ll-13)
Body length: 6.5-7.6mm(from frontmarginofhead toanalend); 3.2-3.5mm
(fromfrontmarginofheadtoelytra1apices).
Body elongate, parallel-sided and somewhat depressedabove. Colour reddish
browntoreddishblackandmoderatelyshining,withmouthparts,antennaeandlegs
brownishyellow.
Maleand female. Thepresentnewspeciesissimilarinbodysizeandgeneral
appearancetoLk〔)yasanum,butdifferentfromitinthefollowingpoints:
Headsomewhattransverse(width/length=1.05)andslightlymorestronglyna一「
rowed anterjad than jn L ko、,asanun1; lateralsidesmoredistinctlyarcuate;Su「face
sljghtlylesscoarselypuncturedthaninL koyasanumandcoveredwithsomewhatf?lIe「
m1croscopjcgroundsculpturethanthatof L koyasanum;eyessmallandnea「lyflat,
thejr1ongjtudjna1djameterlessthanone-thirdaslongaspOSOtCula「Pa「t・Antennae
elongate,extendingtothemiddleofpronotumandnotthickenedOtWa「dStheapical
segment,6thto10thsegmentsmoreorlessmoniliform,andsimilarina「tiCulatiOnto
thoseofL ko、,asanum.
52 Yasuaki WATANABt
Figs. 11-13. MalegenitalorganofLa1/11-oblltm(s.str)o/1to1/e11sesp nov;dorsalview(11).lateralview
(l2),andventralview(13).Scale:1.0mm.
Pronotum similar inconfiguration tothat of L ko、,asamtm, remarkably longer
thanbroad(length/width=1.27),apparently longer(pronotum/head=1.47)andsome-
whatbroader(pronotum/head=1.10) thanhead;surfacesimilarlypuncturedas in L
koyasanlunexcept foranarrowmediansmoothspace.Elytrasubtrapezoida1though
less dilatedposteriadthan inL ko、、,asanum,alittletransverse (width/length=1.10),
distinctly shorter (elytra/pronotum=0.75) but somewhat broader (elytra/prono-
tum=1.05) thanpronotum;posteriormarginbroadlyemarginateatthemiddleasinL
koyasanum;surfacemorecloselyandmoredistinctlypuncturedthaninLko、,aanMn.
Legsmoderatelylong,profemora,protibiaeandprotarsisimilarinstructuretothoseof
L ko、,asamm1.
Abdomenelongate, 3rd to7th tergiteseachmoredenselyandmorecoarsely
punctured than in L ko、,asanun1andcoveredwithfinebrownishpubescence,8thter_
gitedistinctlymoresparinglypuncturedthanintheprecedingtergjles; jnmale,81h
Ste「nifeme「eshallowlyemarginateatthemiddleofposterjormargjnthanjnLk()、,a_
Sanlml andShallowly,nock-shapedlydepressedinfrontoftheemargjnatjon,surfaceof
thedeP「eSSOinmorecloselysettledwithfinebrownishsetaethanjnotherparts;7th
Ste「nifeSli9htIyemarginateatthemiddleofposteriormarginandshallowly,1ong1tud1_
nattydeP「eSSedbeforetheemargination,surfaceofthedepressionmorecoarselyso_
tOSethaninothe「Parts;6thsternitesimple;infemale,8thsterniteabruptlynarrowed
inPOSte「Oi rthirdtowardsthenarrowlyroundedapex;7thsternjtesjmple
Male9enita1organspindle-shapedandwellscIerotizedexcept formembraneous
ApterousLath,・()11tl,11fromtheKiiPeninsula 53
Ventralsideofmedianlobe.Medianlobeconsiderablyshorterthanfusedparamere,
widestatbasalthirdandmorestronglynarrowedapicadthanbasad,withventralscle_
「iteeton9ate, muchshorter thoughapparently broader than that of L koyasartMn,
ab「uptlynarrowedinapicalpart towardsthebluntlypointedapex.Fusedparamere
Slightlyasymmetrical inapical third,abruptlynarrowedinapical thirdtowardsthe
acutelypointedapex;surfaceprovidedwithafinelongitudinalcarinaatthemjddlejn
apical fourth.
Typeseries. Holotype: , allotype: 0sugidani,Mt. 0htoh-san,Wakayama
,
P「ef.,Honshu,Japan,24-IV-1990,1.MAToBAleg.Paratypes:1(;,samelocalityand
Collectorasabove,29-VII-1982;1 Mt. 0htoh-san,WakayamaProf.,Honshu,Japan,
,
15-V-2000, 1. MATOBA leg; 2 Akatsuchimori-yama, 0htoh-mura, Wakayama
,
Prof.,Honshu,Japan,26-V-1993,1.MAToBAleg;1d,Maenokawa,WakayamaProf.,
Honshu,Japan,24-V-1991,1.MAToBAleg.All thetypespecimensaredepositedin
thecollectionoftheLaboratoryofInsectResources,TokyoUniversityofAgriculture.
Distribution. Japan(centralHonshu:KiiPeninsula).
Bionomics. ThetypespecimenswereextractedbyaTu11gren funnel fromleaf-
littertakeninbroadleavedforestsofOhtoh-murainWakayamaPrefecture.
Etymology. Thespecificepithetofthisnewspeciesisgivenafterthetypelocal-
ity“Mt. 0htoh-san”.
Lat1trobium(s.str.)isenseY.WATANABE,spnov.
[Japanesename:Ise-himekobane-nagahanekakushi]
(Figs4,14-16)
Body length:5.6-6.1mm (from frontmarginofheadtoanal end); 2.9-3.0mm
(fromfrontmarginofheadtoelytra1apices).
Similar in faciesandcolour to L. ohtohe11se, but differs from it in the smaller
bodyandthefollowingpoints:
Male and female. Headslightlytransverse(width/length=1.03)oraslongas
broacしsubtrapezoida1, somewhat more strongly narrowed anteriad than in L.
ohtohense; lateralsidesgentlyarcuate;surfaceslightlymorenumerouslythoughless
coarselypuncturedthaninL.ohto1/enseandcoveredwithmuchfinergroundsculpture
thanthatof L. o11tohense; eyessmall,almost at, their longitudinal diameterabout
one_thjrdaslongaspostocularpart.Antennaeelongate,extendingslightlybeyondthe
mjddleofpronotum,notthickenedtowardstheapicalsegmentasinL.ohtoherlse,5th
10thsegmentsmoreorlessmoniliform,andsmi ilarinarticulationtothoseofL.
to
o/1tohense.
pronotumslightlymoredistinctlynarrowedposteriadthaninL.ohto11enSe,aPPa一「
ent1y longer than broad (length/width=1.23), considerably Ion9e「(P「onOtum/
head=137)andsomewhatbroader (pronotum/head=1.08)thanhead; late「alSides
almoststrajght jn anterior two-thirds,somewhatarcuate inpOSte「iO「thi「d;Su「face
slightlymorenumerouslyandsomewhatmorecoarselypuncturedthaninL・ohOthenSe
54 、11suakiWATAN、1]1
Figs.14-16. MalegenitalorganofLat1/1・chiff,11(s.str)1se11sesp nov: dorsal vie、、'(l4). lateral view
(15),andventralview(16).Scale:1.0mm.
except foranarrowmediansmoothspace.ElytrasimilarinconfigurationtothoseofL.
ohtohense,somewhat transverse(width/length=l.11),distinctlyshorter(elytra/prono-
tum=0.79) butalittlebroader (elytra/pronotum=1.08) thanpronotum;surfaceless
closelyandmuchmorestronglypuncturedthaninL. ohtohense. Legssimilarinstruc-
turetothoseofthemembersofthisspecies-group.
Abdomenelongate,eachtergiteclosely, finelypuncturedandcoveredwithfine
brownishpubescenceasinL.ohto/1ense;inmale,8thsterniteshallowlythoughsome-
whatmoredeeplyemarginateatthemiddleofposteriormarginthaninL.ohtohense
andlongitudinally,morestronglydepressedbeforetheemarginationthaninL.ohto/1-
ense,surfaceof thedepressioncloselysettledwithblackishbrowncoarsesetae;7th
sternitenearlytruncateatthemiddleofposteriormarginandslightlydepressedorflat-
tenedin frontofthetruncation;6thsternitesimple; in female,7thand8thsternites
eachsimilarinconfigurationtothatofL.o/1to17e11se.
Malegenitalorganellipticalandalmostsymmetrical,wellsclerotizedexcept for
membraneousventralsideofmedianlobe.Medianlobesomewhatlongeranddislinclly
broaderthanfusedparamere,widestatthemiddleandmuchmorestronglynarrowed
POSte「Ia Withventralscleritestronglynarrowedtowardsthepojnted
apex Fused
Pa「ame「eab「uPtlynarrowedinapicalparttowardsthepointedtip;lateralsjdestrongly
elevatedinPosteriorhalf,surfaceprovidedwithafinelongitudinalcarjnaalongthe
medianline,bearingatriangularprocessbeforetheapexasseenfromlateralside
「jPeSones・ Holotype: , allotype: Tsurugitouge, Nansej T., Mje pref
,
Honshu,Japan,20-I-2000,H.YoKozEKIleg.Paratypes:13
11 , same data as
fo「theholotype; I 1 ,samelocalityandcollectorasabove,,10_l_1gg8; l i
,
Samelocalityandcollectorasabove,15-I-2000;9
1 Kourajbjro, lse c M1e
P「ef・,Honshu,Japan,10-I-1998,H.YoKozEKlleg;4 , ,sam, elocalityandco1jector
AptcrousLat1/1)/)1tl1,1fromtheKiiPeninsula 55
as above, 31-I-1998; 2 2 Erihara, Isobe T., Mie Pref., Honshu, Japan,
, ,
20-I-2000,H.YOKOZEKIleg.Thetypespecimensaredepositedinthecollectionofthe
Laboratoryof InsectResources,TokyoUniversityofAgriculture,except for several
paratypeswhicharepreservedinYoKozEK1'scollection.
ist''10tfffo'7. Japan(centralHonshu:KiiPeninsula).
!o'7o1mcs. Unknown.
Etvn1o1ogy. The specific epithet of thepresent new species is derived from
“Ise”,theoldnameofMiePrefecture,inwhichlieail thethreeknownlocalities.
Latltrobiuln(s.str.)nankienseY.WATANABE,sp
nov.
[Japanesename:Nanki-himekobane-nagahanckakushi]
(Figs5,I7-22)
Bodylength:5.8-7.4mm(from frontmarginofheadtoanal end); 3.1-3.3mm
(fromfrontmarginofheadtoelytraIapices).
Similar ingeneralappearancetotheprecedingthreespecies,butdistinguishable
fromthembysecondarysexual character ofthe8thabdominalsterniteinthemale,
whichismoredeeplyandsubtriangularlyexcisedatthemiddleofposteriormargin.
Male and female. Headsimilar inconfigurationto thatof L. lsensethough
slightly more transverse (width/length=1.05); lateral sides gently arcuate as in L.
1sense;surfaceslightlymoresparinglyandslightlymorecoarselypuncturedthaninL.
isenseandcoveredwithextremelyfinegroundsculptureasinL. isense;eyessmalland
nearlyflat, their longitudinaldiametershorterthanone-thirdthelengthofpostocular
part.Antennaeelongate,extendingslightlybeyondthemiddleofpronotum,somewhat
moreslender thanthoseofL.1sense,twoproximalsegmentspolished,theremainings
opaque,5thto10thsegmentsmoreor lessmoniliform,andsimilar inarticulationto
thoseofL. isense.
Pronotum narrowed posteriad as in L. lsense, apparently longer than broad
(length/width=1.18), distinctly longer (pronotum/head=1.37) and a little broader
(pronotum/head=1.10) than head; lateral sidesmoredistinctly arcuateinposterior
thirdthaninanterior two-thirds;surfacemoresparinglyandmorecoarselypunctured
than inL. i.sense,except foranarrowmediansmoothspace.ElytrasubtrapezoidaI,
slightly more dilated posteriad than in L. lsense, a little transverse (width/
length=1.12),distinctlyshorter(elytra/pronotum=0.79)butsomewhatbroader(elytra/
pronotum=1.05) thanpronotum; lateral sidesalmoststraightexceptnear anterior
andposlerjorangles,posteriormarginslightlymoredeeplyemarginatethan in L.
1sense;surfaceslightlymoreclosely,moreshallowlyandmoreroughlypuncturedthan
jnL.1sense.Legsmoderatelylong,similarinstructuretothoseofL.lsense.
Abdomenelongate,3rdto8thtergiteseachsimilarlypuncturedandpubescentas
jrl」L 1・so,t?,e; jnmale, 8thsternitesubtriangularlyexcisedat themiddleofPOSte「iO「
margjnandlongitudinallydepressedbeforetheexcision,surfaceofthedeP「eSSOin
morecloselysettledwithfineblackishbrownsetaethaninotherPartsexceptfo「ana一「
56 Yasuaki WI、TANABI
Figs.17-19. MalegenitalorganofLa11/,・obi1l111(s.str)nankie,liesp nov.,、lromIkenoyamaofKoza
gawa-choinWakayamaPrefecture;dorsalview(17),lateralview(18),andventralview(19).Scale
1.0mm
rowlongitudinalsmoothspacealongthemedianline;7thsterniteslightlyemarginate
atthemiddleofposteriormarginandfeebly,longitudinallydepressedin frontofthe
emargination;6thsternitesimple;infemale,8thsterniteabruptlynarrowedinposterior
thirdtowardsthenarrowlyroundedapex;7thsternitesimple.
Male genital organ considerably different from thoseof the three preceding
species.Medianlobeelliptical,slightlylonger thanfusedparamere,withventralscle-
ritewidest at themiddleandabruptly narrowed inapical fifth towards thepointed
apex. Fusedparamerenarrowedtowardstheapex which isacutely pointedasseen
fromdorsalside,stronglycurveddorsadinapical fourthinprofile,surfaceprovided
withafinelongitudinalcarinaatthemiddle.
Typeseries. Holotype: (3,allotype: Ikenoyama, Kozagawa-cho,Wakayama
,
Pref., Honsh1L1,Japan, ll-V-1996,1.MAToBAleg. Paratypes:4 ,9 ,samedataas
for theholotype;3 2 ,samelocalityandcollectorasabove,3-XI-2000;3
, ,
9 Fudou-dani,MiyamaT.,MiePref.,HonshLl,Japan,22-IX-2001,K.TOYODAleg;
,
14 10 Mikizaki, 0wase C., Mie Pref., Honshu, Japan, 25_x_1995, H.
, ,
YOKOZEKIleg;2 ,samelocalityandcollectorasabove,29-IX-1996; 13 18
, ,
Kukizaki,0waseC.,MiePref.,Honshu,Japan,12-II-1996,H.YoKozEKl leg;3
Same locality and collector as above, 1-VI-1996. The bolo_andallotypes aredo_,
PositedinthecollectionoftheLaboratoryofInsectResources,TokyoUniversjtyof
A9「iCultu「e. Theparatypesaredistributed tothecollectionof theabovementioned
laboratory,WakayamaPrefecturalMuseumofNaturalHistoryandYoKozEKl,scoffee_
tion.
tut'the「Speel'nonsexamined. 3 1?,Tategasaki,KumanoC,Honshu,Japan,
,
29-XII-2000, H. YOKOZEKI leg; 3 Kasuga-jinja, Kainan_shi, wakayamapref,
,
Honshu, Japan, 15-IV-1994, 1. MAToBA leg; 4 2 , Aisuji, shingu_shj,
,