Table Of ContentGayanaZool. 56 (1-2): 49-57, 1992 ISSN0016-53IX
AEGLA EXPANSA, NEW SPECIES (CRUSTÁCEA:
DECAPODA:ANOMURA: AEGLIDAE), FROM THE LOWER BIO-BIO
RIVER BASIN, CONCEPCIÓN, CHILE.
AEGLA EXPANSA. NUEVA ESPECIE (CRUSTÁCEA: DECAPODA:
ANOMURA: AEGLIDAE), DEL SECTOR INTERIOR DE LA CUENCA
DEL RIO BIO-BIO, CONCEPCIÓN CHILE.
Carlos G. Jara*
ABSTRACT RESUMEN
Aegla expansa, new species (Decupoda: Anomura: Se describe Aegla expansa, nueva especie (Decapoda:
Aeglidae). from a small forest stream in Concepción Anomura: Aeglidae). recolectada en un riachuelo de la
province.Chile, isdescribed. Itsmostsalientfeaturesare: provinciadeConcepción.Chile. Secaracterizaportener:
1) greatly swollen chelae with expanded subcrescentic I) quelas voluminosas, con cresta palmar subdisciforme
palmar crest; 2) earlobe-like expansión at rear end of muy expandida; 2) extremo posteriorde lacresta palmar
palmarcrestfittinginwideembaymentatdistodorsalend extendidaenformade lóbuloauricularcuyobordeencaja
ofcarpas: 3)dorsal surface ofchelae densely covered by enunampliosenosobreelextremodistodorsaldelcarpo;
minute subacute scales: 4) rostrum broad. ligulate, 3) superficie dorsal de las quelas con densa cubierta de
longitudinally excavated both sides of rostral carina; 5) diminutas escamas subagudas; 4) rostro ancho, ligulado,
extraorbitalsinuswide.separatedfromorbitalsinusbytwo excavadolongitudinalmenteaamboscostadosdelacarena
spines; 6) gastric and cardial áreas markedly convex and rostral; 5) seno extraorbitario ancho, separado del seno
protuberant; 7) carapace branchial margins crenulate; 8) extraorbitariopordosespinas;6)áreasgástricaycardíaca
venteroffourththoracicsternum smooth;9)anterolateral marcadamente convexas y protuberantes; 7) márgenes
angleofsecondabdominalepimeronsharplyacute.Because branquialesserrados;8(superficieventraldelcuartoesterno
the shape of the carapace and rostrum it resembles A. torácico lisa; 9) ángulo del segundo epímero abdominal
araucaniensis Jara but the shape of the palmar crest agudo. PorlaformadelcaparazónydelrostroA. expansa
approaches it toA. laevis laevis (Latreille) and A. laevis se asemeja a A. araucaniensis Jara, en tanto que por la
talcahuano Schmitt. The expanded palmar crest of A. forma de la cresta palmar se asemeja a A. laevis laevis
expansa was probably acquired indcpendently from that (Latreille)yA.laevistalcahuano Schmitt. Esmuyproba-
found in the BrazilianA. schmitti Hobbs III. bleque la ampliacrestapalmarhayasidoadquirida inde-
KeywordS:Crustácea.Anomura.Aegla,newspecies.Bío- pendientementeporA. expansayporA.schmittiHobbsIII
de Brasil.
Bíoriver, Chile.
Palabras clavkS: Crustácea. Anomura, Aegla, nueva
especie, río Bío-Bío,Chile.
*InstitutodeZoología. Universidad Austral deChile.
Casilla567,Valdivia,Chile.
49
Gayana Zool. 56( 1-2), 1992
INTRODUCTION DESCRIPTION
Among the about 40 species ofAegla heretofore Carapacealmond-shaped;gastricandcardialáreas
described (see Martin & Abele. 1988), there is a markedly convex and rotundly prominent along
small numberdistinguished by possessing one or midline. Dorsal surface ofcephalothorax and ab-
two very characteristic morphological features. domen finely punctate.
One ofthis ¡s A. schmitti Hobbs III, 1979, from Rostrum long and broad, surpassmg corneae
Curitiba. Brazil, which has a broadly expanded by at least one cornea length, triangular at base,
and deeply excávate subdisciform chelipedal ligulate in the middle and conical at apex; apex
palmar crest, almos! as vvide as the palm proper. recurvedandtippedbyacuteconical scale. Rostral
BecauseofthatHobbs111(1979:987)statedthatA. margins with row of closely packed lenticular
schmitti stands farout from the remaining species scales; at rostral apex marginal scalesbecome fíat
ofAegla. though he mentioned the Brazilian A. and more separated. Apical scale surrounded by
castro Schmitt,A. o. odebrechtii MüllerandA. o. four tiny ones.
paulensis Schmitt. as its morphologically closest Rostral carina markedly prominent. extended
relatives. In this paper a new species ofAegla is from protogastric eminences to base of rostral
described. basedon only one adult male specimen apex; there it merges into rostral body; ridge of
collected in the province of Concepción, Chile, carina with irregular band of lenticular scales;
which also features an enlarged palmar crest. aboutthreerowsofthematthemostelevatedpoint
ofit. one row on distal carina, and irregularpatch
of7 or8 scalesondorsumofrostral apex. On both
Aegla expansa, new species. sides ofcarinadeep and wide trough make rostral
margins appear rimmed.
Figs. la. h. c. d. e. I. g. h. i, k. I. ni. n. o, p. q, r.
Orbital sinusdeep. separatedfromextraorbital
sinusbysmallconicalspine,itsapexturnedlaterad;
HOLOTYPE
below it and slightly displaced to body midline a
similar but smaller spine pointing out
InstitutodeZoología,UniversidadAustraldeChile,
straightforwardly. Extraorbital sinus wide and
IZUA C-761, adult male; Hualqui riverat La Leo-
shallow. Anterolateral spine acute. slightly
nera,aboutókmeastofHualqui(36°58'S,72°56'
W),and24 KmsouthofConcepción, VIII Región. upturned,arisingfromfíatdorsumofanterolateral
Chile; lOfebruary 1987;J. Llanos coll. lobe; this last one rough. its external margin with
irregular row of scales similar to those on rostral
margins.
Diagnosis Gastric área markedly convex and prominent,
most elevated at level of third hepatic lobes.
Rostrum long, broad ligulate, longitudinally Epigastric prominences wedge-shaped (cunéate),
excavated both sides of prominent dorsal carina; most prominent just behind deepest point of
extraorbital sinus wide U-shaped, separated from orbital sinuses; summit of left one crater-like.
orbital sinus by two spines, one above the other; surroundedby8or1)tinyconicalscales;summitof
gastricandcardialáreasmidline markedlycon\ex right one scabrous and tipped by 9 or 10 scales.
and protuberant; carapace branchial margins Protogastric eminences low but well-defined,
crenulate; propodus of chelae greatly swollen; summits tipped by 1 or 2 fíat lenticular scales.
palmarcrestexpandedasdorsallyconcaveearlike Sparse tiny lenticular scales forming broad band
lamina, its rearend fitting in wide embayment of along midline ofgastric área.
distodorsalendofcarpus;dorsal surfaceofchelae Hepatic lobes separated by shallow grooves
denselycoveredbyminutesubacutescales;venter forming noticeable identations at margin of
of fourth thoracic sternum smooth; anterolateral carapace. First hepatic lobe separated from
angleofsecondabdominalepimeronsharplyacute. anterolateral lobe by wide deep notch, its anterof
50
Aeglaexpansa n. sp.: Jara,C.G.
lateral angle tipped by small acute scale not broad-based knob proximally and small conical
particularly more prominent than those forming knobs tipped by scale distally; one distal knob on
irregular scabrous row along hepatic margin. left and three on right. Ventromesial border of
Dorsum of first and second lobes excávate; third merus with one prominent conical spine at distal
slightly swollen. Posteriorgastricpits (see Martin end. and similar but somewhat smaller spine at
andAbele, 1988)scarcelymarked.Cervicalgroove conespondingsiteonexternalventrolateralborder.
well-defined,particularlynarrowanddeepbetween Externalmost point ofdistoexternal articular lobe
posterolateralportionsofgastricáreaandinnermost ofmeruswithsmallsubapicalspine;proximalhalf
part of swollen inner branchial áreas. of dorsal longitudinal ridge straight or at most
Pterygostomial angle tipped by short stout spine. sinuous;distalhalfwithrowofacutesmalltubercles
Cardiac área almost as long as wide. Areola increasing in size distally, three tubercles on the
broad, markedly convex and prominent. summit leftandfiveontherightmerus.Distodorsalmargin
reaching same height as gastric área; frontal end ofmerus with a very low and ill-defined median
subtriangular, distal point raising sharply from tubercle recognizable by short transversal row of
wide central part of cervical groove. carrying threetofourworn-offconical scales;externaltoit
patch of about 10 loosely packed tiny lenticular somewhat scabrous border bearing irregular row
scales. Dorsumofbranchialáreasslopingdownat oftiny scales.
rather pronounced angle to not particularly Catpus massive; carpal crest formed by four
expanded branchial margin; row of tiny cortical clear-cut,conical,slightlyupturnedspinesincreasing
scalesalonglineaaeglicalateralis(seeMartinand insi/edistally;carpallobelong,markedlydisplaced
Abele.op.cit.).Posteriorbranchialmarginslightly towards distodorsal end of carpus. leaving large
upturned. Epibranchial tooth prismatic and concave gap between it and distalmost spine of
moderately prominent. external margin with row carpal crest; carpal lobe low and rounded off.
oftinyconicalscales,thatatsummittwiceas large bearing small subacute scale at summit; on left
as remaining scales. Border ofanterior branchial cheliped tipofit isbroken. Dorsal ridge ofcarpus
areaminutelydenticulate;denticlesdorsoventrally well developed, made up of seven broad based
depressed. separated by shallow notches; each tubercles tipped by a somewhat oblique row of
with irregularrow ofconical scales, anteriormost three to seven bluntconical scales, that at summit
being most prominent. Nine denticles on left side largest. Lateroexternal ridge of carpus almost
andeightonrightside.Borderofposteriorbranchial absent; from its remains one (left) or two (right)
áreawith singlerow oftightly packed tiny scales. fíat tubercles bearing at front row of small
Right posteriorbranchial áreadeformed by bump lenticularscales. Ventralfaceofcarpuswithshort
probably oftraumatic origin. stoutconicaltubercletippedbysmallscaleleaning
Abdominal epimera deeply punctate. Dorsum distally;someshortstiffsetaebetweenventraland
ofsecondexcávate; anterodorsal angle spiniform, crestal spines.
tipped by short conical scale, posteriorly merging Chelae ovoid in outline, bulky and markedly
intoepimeron throughbroad ridge. Anteroventral inflatedatbothdorsal andventralfaces,especially
angle rounded off. Lateral surface of epimeron, left one. Ventroexternal ridge of palm, between
between anterodorsal and anteroventral angles, lateroexternal knob ofcarpo-palmjoint and apex
markedly concave. Angle of third and fourth offixed finger, arcuate, broad and faintly marked
epimera also acutely tipped. In this specimen left onrighthandbutabsentonleftexceptatdistalend
fourth angle shattered. Telson píate dimerous. where it merges into body offixed finger. Palmar
subpentagonal. Anterolateral angles of third ventrocentraláreaswollen,traversedobliquelyby
thoracicsternumassmallcompressedknobs.apex row of four widely separated, fíat, scale-tipped.
of axial portion blunt and flanked by 3 scales. smalltubercles.Fixedfingerrelativelyshort,stout,
Fourth thoracic sternum fíat. deeply concave at ventral side. cutting edge pro-
Chelipeds very large and bulky. left larger. nouncedlyrecurvedü-shaped)whenseenfromabove;
Ventromesial borderofbasi-ischiums with round. marginofpalm-dactylusjointwithrowoftinyscales.
51
)
GayanaZool. 56(1-2), 1992
Dorsumofpalmscoveredbytiny,flat,lenticular
to conical scales which become more prominent
onlateroexternaldistalhalf.Palmarcrestexpanded
asear-shapedlamina,largeronriglithand;dorsum
oflaminamarkedlyconcaveandprofuselystudded
byminutescales,largeratupturnedmargin,forming
a marginal band; earlobe-like rear end of crest
markedly recurved. fitting into wide indentation
betweentipofcarpallobeanddistal spineofcarpa!
crest when chela flexes upon carpus. Ventrally,
palmarcrest arises directly from proximal palmar
border without furrow ortrough separating them;
ventromarginal tace scabrous and scaly. Dactylus
ofchelae noticeably recurved; when chelae cióse
dactylus and fixed fingercontact only at theirtips
leaving wide gap along most of them; dactylus
with sharp spinitorm lobe on proximal end (right
hand) or two to three tiny acute scales in row at
corresponding site (left hand).
Pereiopodsrobust.Distodorsalmarginofmerus
fringed by short stiffsetae intermingled with tiny
indentations; small acute tubercle at midpoint.
Posteroventral margin ofmerus minutely seríate.
Distodorsalendofcarpusacutebutnotprominent.
Distal thirdofventral marginofdactyluswithrow
of 10 to 14 tiny spinitorm slender scales.
MEASUREMENTS:
Thedimensionsofthe holotypearegiven inTable
1. Measurements were made with calipers to the
nearest 0.1 mm. Morphometric characters
considered are those defined by Jara and López
( 1981 );tothose,fiveothers(markedwithasterisk
are added to provide quantitative grounds for
comparison ofA. expansa withA. schmitti Hobbs
Sex
Aeglaexpansa n. sp.: JARA,C.G.
furrows and depressions of the carapace, frontal 1979), but on the other handA. expansa does not
articularfaceofabdominal terga, andventral face appearparticularly relatedto any Chilean species
ofpereiopodsleastthedactylus. Paléorangetored either.AexpansadiffersfromtheBrazilianspecies
orange on ventral face ofcutting edges of chela, mainlyintheshapeofthepalmarcrestandrostrum.
and palm-dactylus articular knob. Dorsum of In A. expansa the palmar crests are subeliptical,
chelipeds with dirty grayish hue upon indefinible comparatively littleprotuberantoutgrowthsofthe
creamy background. posterointernalborderofthepalm,scarcelyforming
an independent unit beyond the palm itself. This
situation is clearly appreciated when viewing the
DlSTRIBUTlON: palm from underneath, because there is notrough
or furrow along the crestal base as is found in A.
Known only from the type locality. a small river schmitti and A. castro. In A. o. odebrechtii the
which drains a seemingly small basin on the ventral face ofthe palmar crest is much the same
northern margin of Bío-Bío river, Concepción as inA. expansa but its rearend does not form an
province, VIII Región. Chile. ear-like lobe. Furthermore,theshapeofthechelae
is different too. In A. expansa the palms are so
inflatedthatthelateroexternalridgeontheirventral
Naturalhistory. side, between the tip of the fixed finger and the
ventral knob ofthe palm-carpusjoint, is scarcely
Nothing is known about the natural history ofA. marked. In contrast, A. o. odebrechtii has three
expansa, and possibly Hule will be known in the well marked ridges grossly punctate: one at the
future asthesmallbasin in which thetype locality verybaseofthepalmarcrest.one.alongthemedian
is located is being drastically altered by forestry lineofthepalmandonebetweenthetipofthefixed
exploitation. A short visit made to the spot on 18 fingerandapointjustinsidetheventralknobofthe
september 1988 allowed to observe a moderately palm-carpusjoint.Thecentralridgeisobsolescent
fast running stream, 4 to 5 meters wide and no in A. prado and A. schmitti. In dorsal view the
more than 1 meterdeep, in anarrow creek. Water chelae of A. expansa are regularly ovoid while
was yellowishbrown in colorand very turbid; the those of A. schmitti and A. odebrechtii are
unstable substrate was a mixture offine sand and subrectangular in outline; those ofA. castro are
coarse gravel rich in mica andquartz. Itevidently almost as ovoid as those of A. expansa but are
had smothered the once stony or rocky bottom markedly dorsoventrally depressed, so that their
with up to40 cm deep layerofloóse material. No lateroexternal border is very narrow while in A.
Aegla specimens were found after two hours of expansa it is rotund blunt. The orientation ofthe
collecting efforts along 200 m stretch ofriver. palmarcrestdiffersbetweenChileanandBrazilian
species as well; inA. expansa the crestextends in
the same horizontal plañe as the rest ofthe chela,
ETYMOLOGY: whileinAschmittiandA.castrothecrestallamina
From expansa (Latin), expanded. Alluding to the is downwardly inclined. so that it forms a wide
angle respect to the horizontal plañe ofthe chela.
broadpalmarcrestofthechelipedsthatcharacterizes
Other differences include the shape ofthe carpal
this species ofAegla.
crest: inA. expansa it is formed by a row ofthree
rather big conical spines which increase in size
COMPARISON: distally and therefore the distal spine, behind the
distal carpal lobe, is the largest; in the Brazilian
On morphological ground A. expansa appears speciesinsteadthecarpalcrestisformedbyfouror
unrelatedtothe BrazilianA. schmitti,A. castro,A. fivealsoconicalspinesbutthe largestintherowis
o. odebrechtii and A. o. paulensis that also have thepredistalone, whilethedistal spine islessthan
greatly expanded palmar crests (see Hobbs III, half robust the precedent and is slightly dorsad
53
GayanaZool.56(1-2), 1992
displaced. At the large embayment of the left surfaceofchelae studded by small elongate fíat to
cheliped ofA. expansa, hetween the distal spine conical scales.
and the carpal lobe, there is a small protuberance A. expansa differs from A. I. laevis and A.
where, in the Brazilian species, the small distal araucaniensis in having a comparatively higher
spine is found. rostralcarinaridgedbyconspicuousbandofscales;
The rostrum of A. expansa appears as the awelldefinedextraorbital sinuslimitedat internal
paradigmofthePacificrostrumtypecharacterized endbyprominentclear-cutorbitalspine;themargin
by Schmitt (1942). The long Hgulate rostrum, of palmar crest entire including the palmar lobe,
elegantly upturned at the acute tip, markedly therefore without right angled notch at distal end
troughed on both sides ofthe pronounced rostral of the crest; and the margin of the palmar crest
carina, and outlined by wide, slightly upturned subcrescentic instead ofsubrectangular.
margins, differs greatly from the comparatively WithA. I. talcahuanoitsharesthesubcrescentic
shorter, steep sloped, scarcely marginated and palmar crest, the lack of a right-angled notch
untroughed rastra ofthe Brazilian species. On the between the distal end ofthe crest and the palmar
other hand, A. expansa resembles ,4. schmitti in lobe.andtheorbital spine.A. expansadiffersfrom
havingaprominentsubrostral process(seeMartin A. I. talcahuano in having a markedly upturned
and Abele, 198S). but the shape of it differs backward extensión ofthe palmarcrest which fits
markedly between both species. in the ampie embayment between the distal spine
The shape of the carapace of A. expansa is of the carpal crest and the distal carpal lobe; a
howeverquitesimilartothatofA.sí limita'andeven clear-cut orbital spine that delimits a U-shaped
more to that ofA. castro. In the last species the extraorbitalsinus;prominentepigastriceminences;
gastric área elevates above expanded and almost raised rostral carina with notorious bandofscales
fíathepatic lobesasitoceursinA. expansaandthe along its ridge; and flattened hepatic lobes and
cardiacáreahaswideprotuberantareola,similarto anterolateral lobe ofcarapace.
that oftheChilean species. Theydifferbecause in The particular delimitation of the orbit and
A. expansa the frontal end of the areola raises extraorbital sinus by a pairofdistinct spines, one
abruptly above a wide depression of the central abovetheother,isalsofoundinA. rostrataJaraand
partofthecervical groove whichdoesnotexistsin A. bahamondei Jara, but apart from this feature
anyofthe Brazilian species. Anotherdifference is thereseemstobenoothersincommonbetweenthe
themicrodenticulationA.expansahasatthemargin above mentioned species and A. expansa.
oftheanteriorbranchialárea,whichresemblesthe
larger denticles present in the Chilean species A.
denticulata Nicolet and A. bahamondei Jara, and REMARKS
which Brazilian species lack.
A. expansa partially shares morphological Thegeneral áreaoftheNahuelbutaCordillera,and
featureswithatleastthreeChileanaeglids,namely, in particular the área around Concepción city.
A. laevis laevis Latreille, A. laevis talcahuano seemstobea"hotspot"intheevolutionaryhistory
Schmitt. andA. araucaniensis Jara. With the first oftheChilean Aeglids. Infact,atleastfourspecies
and third species it shares the almond-shaped andonesubspeciesnavebeenpreviouslyrecorded
carapaceandligulaterostrum.Thislastparticularly in the territorial stretch between Imperial to the
similartothatin A.araucaniensis. Withbothlaevis south and Talcahuano to the north (see Schmitt
it shares a low blunt carpal lobe. globose hulky 1942.Jara 1982. 1986).Ofthese,three,namelyA.
chelaeandexpandedpalmarcrest. Withall three it concepcionensis, A. I. talcahuano and A. papudo
shares fíat fourth thoracic sternum, acute have been recorded from nearby Concepción city
anterolateralangleofsecondabdominalepimeron. and to themA expansa is nowadded.The reasons
surface ofcarapace distinctly punctate, tubercles forsuchahighdiversityofaeglidsinthatrestricted
oncarpaldorsalridgeelongatenoduloseandtipped geographical áreaare lar from being clearbut it is
by an oblique row offíat lenticularscales. and the interestine to note that none of the records have
54
Aegla expansa n. sp.: Jara,C.G.
beenmadefromtheBío-Bíoriver(seeBahamonde Curator of Invertebrates at the Museum of
andLópez, 1963).themain freshwaterbody inthe Comparative Zoology. Harvard University,
área.Unfortunatelythesamplingeffortforaeglids Cambridge,forherreadinesstolendmetheneotype
A
in the área of Concepción has been meager and and paratypes of /. laevis and the holotype and
therefore it is premature to advance any paratypesofA. /. talcahuano, andDr. RaymondB.
zoogeographical hypothesis. Moreover. the Manning, Nat. Mus. Nat. Hist., Washington, his
profound environmental alterations induced by comments to an earlier versión of the MS. This
the human population in the área ofTalcahuano publication was partially financed by grants
andConcepciónmakeimprobablethedevelopment FONDECYT 91-0900 and DID S-91-4, Universidad
ofan accurate pictureoftheoriginal diversity and Austral de Chile.
distributionofaeglids. However.fromthefactthat
A
atleastA.concepcionensisand expansaarefound
L1TERATURE C1TED
insmallmoderatelyfastrunningstreamsthatdrain
the mountanious outcrops to the south of Con-
Bahamonde. N. and M.T. López 1963. Decápodos de
cepción city. I tentatively conclude that they aguascontinentalesenChile.InvestigacionesZoológi-
representlocal highlyderivedspeciesfromstocks
casChilenas 10:123-149,
ofmoreampiedistribution.InthecaseofAexpansa Hobbs. H.H., III. 1979. A new species of the endemic
suchancestralstockcouldhavebeentheforerunners South American genus Aegla from Paraná. Brazil
ofA.araucaniensisand/orA. laevis(sensulato), to (Crustácea: Anomura: Aeglidae). Proceedings of the
which A. expansa seems more closely related. Biológica! Society ofWashington 91(4): 982-988.
Jara. C.G. 1982. Aegla bahamondei, new species
(Crustácea: Decapoda: Anomura) from the Coastal
Mountain Range of Nahuelbuta, Chile. Journal of
ACKNOWLEDGMENTS Crustacean Biology. 2(2): 232-238.
Jara, C.G. 1986. Aegla spectabilis, a new species of
freshwatercrabfromtheeasternslopeoftheNahuelbuta
TheauthorthanksAMr.J.Llanosthedonationofthe CoastalCordillera.Chile. ProceedingsoftheBiological
type specimen of expansa; Dr. Joel W. Martin. Society ofWashington 99(1 ): 34-41.
Natural History Museum ofLos Angeles County, Jara, C.G. and M.T. López. 1981. A new species of
foraphotocopy ofH. H. Hobbs III' description of freshwatercrab(Crustácea:Anomura: Aeglidae)from
A schmitti; the Smithsonian Institution (Was- insular South Chile. Proceedings of the Biological
mhiengttoocn.omDp.aC.re)tthheetheonlwoeteykpefeolflAowesxhippanthsaatwailtlhowtehde MartoSifoncti.heeJt.ygWeo.nfauWsnadAseLh.giGln.agAt(boCenrlu9es4t.(ác11e9)a8::88.8A-En9xo3t.emrunraal:moArepglhiodlaoeg)y.
A
type-series of schmitti, A. 1. laevis and A. I. Smithsonian Contributions toZoology453: 1-46.
talcahuano. and with paratypes ofA. castro, A. o. Schmitt.W.L. 1942.ThespeciesofAegla,endemicSouth
odebrechtii and A. o. paulensis, at the National Americanfresh-watercrustaceans.Proceedingsofthe
MuseumofNaturalHistory;Dr.ArdisB.Johnston, United States National Museum 91(3132): 431-520.
55
GayanaZool. 56(1-2), 1992
Figure \.Aeglaexpansa, holotype.a,dorsal aspect;b,dorsumofprecervicalcarapace;c, lateral profileofthe same;d.
scaleson marginofthe anterolateral lobeofcarapace.orbital spinesandrostrum, left hall'ofcarapace;e, marginofleft
halfofprecervicalcarapaceshowinghepaticlobesandepibranchialtooth;f.scalesonmarginofleftanteriorbranchialárea
andepibranchial tooth.
56
Aeglaexpansa n. sp.: Jara,C.G.
Figure 1.A. expansa, holotype. g,dorsal viewofleftchela;h,sameofrightchela;i,sagitalviewofleftchelipedpalmar
crest;k,ventralviewofrightchelipedpalmarcrest;1,sagitalviewofrightchelipedpalmarcrest;ni,dorsumofleftcheliped
merus; n, ventral view ofproximal anielesofleftcheliped.
Figure \.A.expansa, holotype.o,lateralviewofsecondabdominalepimeron;p,telsonpíate;q,ventral viewofthirdand
fourth thoracic sterna; r, merus. carpus and propodusofleft second pereopod.
57
