Table Of ContentElytra, Tokyo,23(1):77-88,May15,1995
Tax-onomicandFaunisticContributionstotheKnowledge
ofPalaearcticQuediina
(Coleoptera,Staphylinidae,Staphylinini)
AlesSMETANA
Centre forLandandBiologicalResourcesResearch,
BiologicalResourcesDivision,AgricultureCanada,
Ottawa,OntarioKIAOC6,Canada
Abstract TaxonomicandfaunisticdataonPalaearcticQuediina,mainlyon the
speciesof thegenusQuedius,areprovided.Quedius(Microsau・,us)kalabiisdescribedand
加ustratedasnew fromspecimensfromKyrgyzstan(TianShan); Quedius(Quedionuchus)
ar,mpesSHARP,1889isredescribedandillustratedfromspecimensfromKunashirIsland;
Quedius (Distichalius)1oebli SMETANA, I978isplacedinsynonymywith Q. fagelianus
CoIFFAIT, 1967 (new synonymy); Quedius lRaphirus) sparsutus FAUvEL, 1875 and Q.
(Raphirus) lederlBERNHAUER,1902areplacedinsynonymywithQ.sublimbatusMAKLIN,
l853(newsynonymies); sexualcharacters, bothmaleand female, aredescribedand il-
lustratedforQuedius(.Rapflirus)walteriKoRGE,1971fromnorth-easternAnatolia.
Thepresentpaper isthefirstofaseriesofpapersintendedtopresent resultsof
myrecentstudiesonthePalaearcticmembersofthesubtribeQuediina.
Thespecimensmentionedin thepaper aredepositedinseveralcollectionsand
theabbreviationsusedinthetext,whenreferringtothesecollections,areasfollows:
ASCC(A.Smetanacollection,Ottawa,Canada)
NHNG(Museumd'Histoirenature11e,Geneve,Switzerland)
MKCC(M. Kociancollection,Praha,CzechRepublic)
Quedius(Microsaurus)kalabisp
nov.
(Figs.1-6)
Entirely, includingappendages, rufo-testaceous,elytraandabdomenbecoming
inconspicuouslypalertowardapex.Headofobtuselyquadrangularshape,aboutas
longaswide,parallel-sidedbehindeyes,posterioranglesbroadlyarcuate;eyesvery
smalland flat,not protruding from lateralcontoursofhead, temperaabout2.5as
longaseyesseen fromabove(ratio2.46);noadditionalsetiferouspuncturesbetween
anterior frontal punctures; posterior frontalpunctureand temporal punctureboth
situatedmuchclosertoposteriormarginofheadthantoposteriormarginofeye; two
smallsetiferouspuncturesbetweenposterior frontalpunctureandposteriormargin
78 AlesSMETANA
ofhead;temperawithsomefinepuncturesposteriorly;surfaceofheadwithveryfine
and densemicrosculpture ofmostly transverse waves and with scattered micro-
punctulation.Antennarathershort,segment3distinctlylonger thansegment2,seg-
ments4and5aslongaswide,segments6-10wider thanlong,graduallybecoming
shorterandwider,segment10distinctlytransverse,lastsegmentaboutaslongastwo
precedingsegmentscombined.Pronotumslightlywider thanlong(ratio1.14),basal
marginbroadlyrounded, lateral marginsdistinctly,slightlyarcuatelynarrowedan-
teriad; dorsal rows eachwith two finepunctures;sublateral rowseach with two
punctures,posteriorpuncturesituatedslightlybeforelevelof largelateralpuncture;
surfaceofpronotumwithmicrosculpturesimilar tothatofhead,micropunctulation
indistincttoentirelyobsolete.Scutellumimpunctate,surfacewithverydenseandfine
microsculptureoftransverselines.Elytrashort,atbasenarrowerthanpronotumat
widestpoint, inconspicuouslywidenedposteriad,atsuturedistinctly(ratio128),at
sidesslightly(ratio1.12)shorterthanpronotumatmidline;punctationfine,moderate-
lydense,vaguelyasperate,transverseinterspacesbetweenpuncturesmostlyabout2.5
aslargeasdiametersofpunctures;pubescencebrownish;surfacebetweenpunctures
withveryfineandratherdensemicropunctulation.Wingsreducedtonon-functional
stumps.Abdomenwithtergite7(fifthvisible)lackingwhitishapical fringeofpalisade
setae;punctationandpubescenceof tergitesslightlyfiner anddenser than thaton
elytra,punctationsimple,evenlycoveringtergites,graduallybecomingvaguelysparser
towardapexofabdomen;surfacebetweenpunctureswithexceedinglyfineanddense
microsculptureoftransversestriae.
Male. First foursegmentsof front tarsusstronglydilated,sub-bilobed,each
denselycoveredwithmodifiedpalesetaeventrally;segment twovaguelywider than
apexoftibia(ratio1.10);segment fournarrower thanprecedingsegments.Sternite8
with two strongsetae at each side; with moderately wide, shallow, subangulate
medic-apicalemargination,small triangularareabeforeemargination attened(Fig.
1).Genitalsegmentwithtergitel0stronglynarrowedtowardsubtruncateapex,with
twostrongapicalsetaeandwithnumerousfiner,variablylongsetaeonapicalportion
(Fig 2);sternite9subemarginateapically,denselysetose,with threestronger lateral
setaeinfrontofapicalmargin(Fig 3).Aedoeagus(Figs.4-5)small;medianlobewith
shortapicalportionwithnarrowlyarcuatetoobtuselytriangularapexbearingminute
toothonfaceadjacenttoparamerejustbelowapex.Paramerelarge,coveringmostof
medianlobe,graduallydilatedanteriadandthennarrowedintonarrowlyemarginate
apexnotquitereachingapexofmedianlobe; twofineapicalsetaeoneachsideof
medial emargination, two similar setae at each lateral margin quite below apex;
undersideof paramerewithoneirregulargroupofsensorypegsetaeclose to apical
marginoneachsideofmedialemargination,eachwithsevenoreightpegsetae,and
withoneirregular, lateral,groupofsixsensorypegsetaeoneachsidearoundwidest
pointofparamere. Internalsacwithout largersclerotizedstructures.
Female. First foursegmentsoffronttarsussimilar tothoseofmale,butdis-
tinctly less dilated and with less numerousmodified palesetae ventrally; segment
TaxonomyandFaunisticsofPalaearcticQuediina 79
twovaguelynarrower thanapexoftibia(ratio0.92).Genitalsegmentwith tergite10
short and wide, distinctly pigmented medic-apically, markedly narrowed toward
subacuteapex,withnumerousunequallylongsetaeatanteriormarginandonapical
portion(Fig 6).
Length8.5-9.3mm.
Typematerial. Holotype(male): [Kyrgyzstan]: “SU-TienShan NEpart of
Terskey Ala Too ridge3000-3600m”. In the collection of the Naturhistorisches
Museum,Wien,Austria.
Allotype(female):[Kyrgyzstan]: ''SU-Tien-SchanMts,-3000-3500mNOpart
of TerskeyAlaTooridgeJETY-0GUZ,-20kmSWFromPrzewalskJ. Kalableg.
23.-30. 6. 1989”/“QUEDIUS(Microsaurus)sp ”. In theSmetanacollection,
n
Ottawa, Canada.
Paratype (male): same data as holotype. In theSmetana collection, Ottawa,
Canada.
Geographicaldistribution. Quediuskalabiisatpresentknownonly fromthetype
localityon thenorthwesternslopesoftheTerskey Alatau Khrebet oftheTianShan
massivein Kyrgyzstan.
Bionomics. Nothingisknownabout thecollection circumstancesof thespec-
imens.Basedon thepalegeneralcolorationofboth thebodyand theappendages,
andon theverysmallsizeof theeyes, itseemsprobablethat Q kalabi occurs in
subterraneanburrowsofsomeanimal, probablyarodent.
Recognitionandcomparisons. Quediuskalabimayberathereasilyrecognizedby
the combination of both the external characters, and the male and female sexual
characters.ItistosomeextentsimilartoQ duiSMETANA,1989fromPunjab(Himacha1
Pradesh)withwhichitsharesthepositionsofboththeposterior frontalandtemporal
punctures,andthegeneralshapeoftheaedoeagus;however,Qduidiffers,inaddition
tothemalesexualcharacters, includingthedifferenceson themedianlobeandon
theparamereoftheaedoeagus(seefigs 25and27inSMETANA, l989,andFigs.4-5),
bythepresenceofonlytwopuncturesineachof thedorsal rowsonthepronotum,
bythestouter formanddarkercoloration,etc.
Etymology. Patronymic. Thespecieswas named in honor of Mr. J. KALAB,
Jinacovice,CzechRepublic,whocollectedtheallotype.
Quedlus(Microsaurus) tenelius(GRAvENHoRsT)
SlaphylinusteneliusGRAvENHoRsT,1806,54.
Quediustene11us:CoIFFAIT,1978,168.
New records. [Rossia]: Buryat rep. Baikal. Bolsaja Ceremsana, 460m, 20~
25-VII-92, M. TRYzNA(ASCC, MKCC) 2.
Comments. Thisisanorthernspecies,widelydistributed from Fennoskandia
eastwardthroughSiberiatotheLakeBaikalareaandtonorthernMongolia(SMETANA,
1967, 215).
80 AleSySMETANA
Quedills(Distichallus) fagelianusCoIFFAIT
Quedius(Mlcrosaurus)faigelianusCoIFFAIT,1967,391;1978,180.
Quedius(Distichalius)1oeb11SMETANA,1978,121(sy'1.nov).
New record. [Lebanon]: Hasroun pres de Becharre, l,500m, 3-IV-75, C.
BEsUcHET(MHNG)1.
Comments. QuediusfagelianusisatpresentknownfromLebanonandIsrael.
Co1FFAIT(1.c)assigned thisspeciestothesubgenusMicrosaurus.Hesuggested
thatitisrelatedtothespeciesofthe“groupedeQ.lateralis'',particularlytoQfissus,
basedonthefactthattheapicalportionofthemedianlobeinQ ftssusisalsodivided
into twobranches, althoughnot as deeply as in Q fatgelianus. Subsequently, he
establishedaseparatemonotypic“groupedeQ fagelianus'' for it,withinMicrosaurus
(COIFFAIT, 1978,180).
Thecharacterstateofthedividedapicalportionofthemedianlobeapparently
developedindependentlyat least twicewithinQuedius. Itappearsrarely,except it is
frequentinthespeciesofsomeAsiaticspecies-groupsofthesubgenusDistichalius(see
SMETANA,1995).IhavelittledoubtthatQ fagelianusbelongstothetaxonthatisat
presentrecognizedasthesubgenusDistichalius,basedbothontheexternalcharacters
(elytra1punctation)and theapicallydividedmedian lobeoftheaedoeagus.
I havenotstudiedthetypespecimensofQ fagelianus; however, thereis little
doubt that thenameQ.1oebli isajuniorsynonymofQ fagelianus(newsynonymy).
Quedius(Quedl'onuchus)armipes(SHARP)
QuedionuchusarmipesSHARP,1889,34.
Quediusarmipes:BERNHAUER&SCHUBERT,1916,419;ADAcHI,1957,179.
New records. [Rossia]: Kunashir, three different localities in the vicinity of
Mendeleevo,13-IX-72, 18-IV- and22-V-77,PoToTsKAYA(ASCC)3.
Comments. Thespecieshasneverbeenadequatelydescribed;thereforeadetailed
description isgivenbelow.
Description. Piceouswithblackhead, apicalmarginsofabdominal tergitesand
apexofabdomenusuallyinconspicuouslypaler;maxillaryandlabialpalpi testaceo-
brunneous,antennaebrunneo-piceousorbrunneouswithfirst threesegmentsdark-
ened;legsbrunneouswithpalertarsi,medial facesofmiddleandhind femorapiceous-
black. Headofrounded quadrangular shape,distinctlywider than long(ratio130),
Fig. 1-17. - 1-6.Quediuskalabi:1,apicalportionofmalesternite8;2,tergite10ofmalegenital
segment;3,stemite9ofmalegenitalsegment;4,aedoeagus,ventralview;5,apicalportionof
undersideofparamere;6,tergite l0offemalegenitalsegment. - 7-11. Quediusa・,,mpes:7,
apical portionofmalesternite8;8, tergite10ofmalegenitalsegment;9,stemite9ofmale
genitalsegment;10,aedoeagus,ventralview;11,apicalportionofundersideofparamere. -
12-17. Quedius、valteri: 12, apical portionofmalesternite8;13, tergite10of malegenital
segment;14,stemite9ofmalegenitalsegment;15,aedoeagus,ventralview;16,apicalportion
ofundersideofparamere;17,tergite10offemalegenitalsegment.
TaxonomyandFaunisticsofPalaearcticQuediina 81
82 AlesSMETANA
slightlynarrowedbehindeyes,posterioranglesobtuselyrounded;withdistincttrans-
verseimpressionbetweeneyes;eyessmall, onlyslightlyconvex; temperamarkedly
longer thaneyesseen from above (ratio 133); noadditional setiferouspunctures
betweenanterior frontalpunctures;posterior frontalpuncturesituatedclosetopos-
teriormargin ofhead, twoadditionalsetiferouspuncturesmediadof it; temporal
puncturesituatedsomewhatcloser toposteriormarginofhead than to posteriO「
marginofeye,oneadditionalsetiferouspuncturebetweenitandposteriormar9inof
eye;surfaceofheadwithveryfineanddensemicrosculptureoftransversestriae,with
interspersedmicropunctulation. Antennashort,segment 3somewhat longer than
segment2,segment4aboutaslongaswide,segment5slightly,segments6-10distinctly
transverse, lastsegmentshorter than twoprecedingsegmentscombined.Scutellum
impunctate,withextremelyfinemicrosculptureof transversestriaewith intermixed
micropunctulation.Pronotumslightlywiderthanlong(ratio1.10),widest atabout
mjddle,moderatelyroundedbasally,onlyslightlytransverselyconvex,aboutequally
narrowedbothanteriadandposteriad,lateralmarginseach attenedtoinconspicuously
concaveposteriorly;dorsalrowseachwiththreepunctures;sublatera1rowseachwith
onlyonepuncturenearanteriormargin;microsculptureonpronotumsimilartothat
onheadbutinterspersedmicropunctulationfiner.Elytramoderatelylong,flat,atbase
aboutaswideaspronotumatwidestpoint,hardlywidenedposteriad,atsutureslightly
(ratio120),atsidesdistinctly (ratio130) longer thanpronotumatmidline. Each
elytronwithpunctationlimitedtotwoirregular,longitudinal rowsofPunctures,one
closetosuture,otherataboutlateralthird;withslightlongitudinal impressionalong
suture;withseveralspine-likesetaeonhumerusandwithonehumeralandtwolateral
longsetae;surfaceslightly, irregularly wrinkled, with dense and fine, submeshed
microsculpturegraduallybecomingmoresuperficial towardelytra1apex.Wingsfully
developed.Abdomenwithtergite7(fifthvisible)bearingwhitishapicalseamofpali-
sadefrjnge;punctationofabdominaltergitesfineandmoderatelydense,becoming
somewhatsparsertowardapexofeachtergiteandingeneraltowardapexofabdomen,
firstvisibletergitewithsparseandextremelyfinepunctationonmiddlearea;pubes-
cencepiceous;surfacebetweenpunctureswithexceedingly denseand fine micro-
sculptureoftransversestriae.Ventro-media1marginofhindfemoraexpanded,except
nearapex,expandedportionwithnumerous,strong,spine-likeblacksetae.
Male. First foursegmentsof fronttarsusmoderatelydilated,subbilobed,each
denselycoveredwithmodifiedpalesetaeventrally;segmenttwonarrowerthanapexof
tibia(ratio0.80);segment foursomewhatnarrowerthanprecedingsegments.Sternite
8withtwolongsetaeoneachside;withrathernarrowandshallow,arcuatemedic-
apicalemargination,small triangularareabeforeemargination attened and smooth
(Fjg 7). Genital segmentwithtergite10small,strongly narrowed toward arcuate
apex,withtwolongandseveralshortersetaeatapicalmargin,andwithseveralsmall
setaejn front ofthem(Fig 8);sternite9shortandwide,witharcuateapex,densely
setoseandwithtwominuteapicalsetae(Fig 9).Aedoeagus(Figs.10-11)small,short
andwide;medianlobewithapexsubarcuate,withtwolatero-apica1sclerites,apical
TaxonomyandFaunisticsofPalaearcticQuediina 83
portionwithsmall toothon faceadjacent toparamere.Paramerenotquitereaching
apexofmedianlobe,narrow,subpara11e1-sidedwitharcuateapex; four finesetaeat
apexand twosimilarsetaeateachlateralmarginbelowapex;sensorypegsetaeon
undersideofparamerearranged intotwoirregular apicalgroups,eachwithsix or
sevenpegsetae. Internalsacsimple,without largersclerotizedstructures.
Female. Firstfoursegmentsoffronttarsussimilartothoseofmale,butslightly
lessdilated.
Tergite10ofthegenitalsegmentwasseverelydamagedintheonlyfemaleavail-
able; it cannot bedescribed or illustrated.
Length7.9-8.8mm.
Geographicaldistribution. Quediusarmlpe.sisatpresentknownfromtheJapanese
IslandsofKyushuandHonshu(SHIBATA,1984,130)andfromKunashir,thesouthern-
most island of the Kuri1Islands.
Bionomics. Collectioncircumstancesof thespecimensarenot known, but it is
assumed that theyweretaken fromunderthebarkoftrees
)i “edl'“s(Kap詰'rMs walter KoRG
(Figs.12-17)
Quedius、v,alteriKoRGE,1971,44;CoIFFAIT,1978,182.
New records. [Turcia]: “Turk.-Rize7. '76, Valleed.1.Firtina1400m; Vit lgt ”
(ACSS) l;Borcka,BalikliDagi, l4-VI-92, M.JANATA(ASCC, MKCC)2.
Comments. Thespecieswasuntilnowknownonlyfromthetwofemalespeci-
mens, collected at Ilica in the valley of Ardesen (Kackar-Daglari). The two new
recordsseem to indicate that thespecies iswidely distributed in themountainsof
northeastern Anatolia.
KoRGE(1971,44)discussedthedifficultyhehadwiththesubgenericassignment
ofthisspecies.HeeventuallychosetoplaceittentativelyinthesubgenusMlcrosaurus,
butpointedout that itmayactuallybelongtothesubgenusSauridus(recentlycon-
sideredasajuniorsynonymof Raphirus -seee.g.SMETANA,1988, 183).Both the
charactersontheaedoeagusandtheexternalcharacters,particularlythepresenceof
onlyoneadditional setiferouspuncturebetweentheposterior frontal punctureand
theposteriormarginof thehead(seeSMETANA,1988,183) leavelittledoubt that
.
walteribelongstothesubgenusRaphirus(seeabove).
ThespecieswasdescribedindetailbyKoRGE(1971,44)andthereforeonlythe
maleandfemalesexualcharacters,includingthoseonthegenitalsegments,areadded
here.
Male. First foursegmentsof front tarsusstronglydilated, sub-bilobed, each
verydenselycoveredwithmodifiedpalesetaeventrally;segment twoaboutaswide
asapexoftibia;segmentfournarrowerthanprecedingsegments.Sternite8withwide
anddeep,obtuselytriangularmedic-apicalemargination,smalltriangularareabefore
emarginationslightlyflattenedandsmooth(Fig.12).Genitalsegmentwithtergite10
84 AlesSMETANA
fairlynarrowandelongate,narrowlyarcuateapically,withtwostrongerapicalsetae,
andwithadditionalweakersetaeatapicalmarginandonapicalportion(Fi9. 13);
sternite9asinFig.14,withsubarcuateapexbearingtwoapicalsetae.AedoeaguS
(Figs.15_l6)verynarrowandelongate;medianlobenarrowedintosubacuteapex,
withsmalltoothjustbelowapexonfaceadjacenttoparamere;paramerenarrowand
veryelongate,almostparallel-sidedwithnarrowlyobtuseapex,slightlyexceedingapex
ofmedjan1obe,narrowerthanmedianlobeexceptcoveringveryapexofmedianlobe
anteriorly; fourextremelysmallsetaeatapexand twosimilarsetaeat eachlateral
marginbelowapex;undersideofparamerewithoutsensorypegsetae,butwith two
short,longitudinalmedialcarinaejustbelowapex;internalsacsimple,withoutlarger
sclerotizedstructures.
Female. First four segments of front tarsussimilar to thoseof male, but
somewhat lessdilated;segment twoslightlynarrowerthanapexoftibia(ratio0.85).
Genitalsegmentwithtergite10quitedistinctive,narrowedtowardnarrowlyarcuate
apex, with middle portion longitudinally elevated, elevated portion with two fine
longitudinalcarinaeatbase;narrowmedic-apicalportionpigmented;withfiveapical
setaeandwithsomeadditionalsetaeonapicalportion(Fig.17).
Theconfigurationofboththeapicalportionoftheparamereoftheaedoeagus
(lackingsensorypegsetae,but bearingtwolongitudinal carinae) and tergite10of
the femalegenital segment (middleportionelevatedandprovidedwith twobasal
longitudinalcarinae)areuniquefeaturesofthisspecies
l‘ed1lls( l'l1'1s) transsylyaM'eMsWEISE
ap
QuediustranssylvanleusWEIsE,1875,356;SM、ITANA,1962,139.
New record. [Ukraina]:Zakarpatskijreg.,Kvasyenv.,23-VII-92,0.HOvORKA
(MKCC)1.
Comments. Thisisanendemicspeciesof thenortheasternCarpathians,partic-
ularlyoftheCzernahoramassive. 0nlyveryfewrecent recordsareknown
s) Medl1'1s(Kap加'r gs nevu EPPEL田S '
QuediusnovusEppELsHEIM,1892,331;CoIFFAIT,1963,389.
.
New records. [Uzbekistan]:Zeravanskijchr.,AmanKutan,1,250m,9-VI-89,
0.HovoRKA(ASCC)1;Zaamin.zapovednik,15-VI-90,J.MAREK(ASCC.MKCC)6.
Comments. Quediusnoviisseemstobewidely distributed inthemountainous
areasofCentralAsia,butveryfewrecent recordsareknownatpresent
) lledl'lrs( ap加ﾍ Is s“aZl' atMs MAKLIN
Quediussut)1imbatusMAKLIN,1853,190;SMETANA,1971,219.
QuedjussparsutusFAUvEL,1875,XXXIV;GRIDELLI,1922,124;1924,124;C0lFFAIT,1978,250(Syn nov)
TaxonomyandFaunisticsofPalaearcticQuediina 85
Quediuslede''1BERNHAUER,1902,699;GRIDELLl,1922,129;1924,124(necColFFAIT,1967)(synnov).
QuediusarcttcusMUNSTER,1920,57;SMETANA,1965,47.
QuediuSWuO「entausiBERNHAUER,1927,96;SMETANA,1971,220.
New record. [Rossia]:Buryat rep.Baikal.BolsajaCeremsana,460m,20~25_
VII-92,M.TRYzNA(MKCC)1.
Comments. This is acircumpolar species, widely distributed throughout the
northern portions ofEurasiaand North America (seeSMETANA, 1967, 217 for a
distributionalmap).
Dueto thiswidedistribution, thespecies was namedmany times, based
on
specimensfromboth thePalaearcticandNearcticRegions.Someof thesynonymy
waspresentedalreadybySMETANA(1965,47,Q arcticus;1971,220, Q.wuorentausj
andsomeadditionalnamesbasedonNearcticspecimens),butColFFAIT(1967,417;
1978, 205)still recognizedQ arcticusasavalidspeciesandneverevenmentionedQ.
sublimbatus.
Quediussparsutus. FAuvEL (1975, XXXIV) described thespecies fromspeci-
mensfromtheLakeBaikalarea(“RegionduBaikal,Irkutsk,onju11ietetseptembre
(Radae)''. I havenot seen theoriginal specimens. However, based on ColFFAIT's
redescriptionand illustrationsof theaedoeagus(1978,115, fig 46D-F;250)ofthe
species(COIFFAITusedFAUvEL'soriginalspecimensfortheredescription),thereisno
seriousdoubtthatthenameQ.sparsutusisajuniorsynonymofQ. sublimbatus.
Quedius lederi. BERNHAUER(1902, 699) described thespecies fromasingle
specimenfromtheLakeBaikalarea(“einimBaikalgebieteaufgefundenesExemplar
(Bang-Haas)'').ThefemaleholotypeattheFieldMuseumofNaturalHistory,Chicago,
is labelled as follows: dark blue square label/“Baikal Bang-Haas''/“Lederi
Rtt''/“LederiBrh.1eg. LederType''/''ChicagoNHMusM.BernhauerCollection''.
Thespecimendoesnotspecificallydiffer fromthespecimensofQ.sublimbatus; the
nameQ.lederiisajuniorsynonymofQ.sublimbatus.Mycorrespondingdetermination
labelhasbeenattachedtotheholotype.
Thereseemstobesomeconfusionconcerningtheidentityoftheholotypeof
.
1ederi.GRIDELLI(1924,124)citedunderQ.sparsutus''LagoBaica1,Tunkun,Sajan
(Bang-Haas); unesemplare comunicatomidaBernhauer, (tipodel Lederi)''. The
holotype(seeabove) isactuallyafemale,withdifferent labels,sothespecimensent
byBERNHAUERtoGRIDELLIcouldnothavebeentheactual ''type''.Butsinceboth
specimensobviously belong to thesame taxon, theconfusiondoes not affect the
nomenclature.COIFFAIT(1978,205)repeatsthedatagivenbyGRIDELLl (1.c)asthe
typelocalityforQ. lederi inhissense(=Q.Jenisseensis,seebelow).
uedlMS(Kap irMs)jeMsseensis(J.SAHLBERG)
RaphirusJ'enlsseensisJ.SAHLBERG,1880,72.
QuediusJenlsseensis:GRIDELLI,1924,167;SMETANA,1976,24.
Quediuslederi:COIFFAIT,1967,417; l978,205(neeBERNHAUER,1902,699)
86 AlesSMETANA
Newrecords. [Rossia]:Buryatrep.Baikal.BolsajaCeremsana,460m,20~25-
vIl_g2,MTRYzNA(ASCC,MKCC)8;PrimorskijKraj,5kmEK「aSkino,13~16-
VII_92,D.BoUKAL(ASCC,MKCC)3.
commeKts uediusJenlsseensls was until now known only from no「the「n
sjberja, fromabouttheKaninPeninsulainthewest totheLenariverbasininthe
east(SMETANA,1976,25).Thenewrecordspresented,extendthedist「ibutiOna1「an9e
considerablysouthward.TherecordfromnearKraskinoisofparticular interest,since
the1ocaljtyisin theextremesouthwesternportionof thePrimorskij Kraj,nearthe
border with Chinaand North Korea.
ColFFAIT(1967,417;1978,205)redescribedthespeciesunderthenameQ. lederi
BERNHAUER,1902.HestatedthatQ.lederiisaspeciesquitedistinctf「omQ.Spa「SutuS
FAuvEL,1875andnotasynonymofthelatter,asproposedbyGRIDELLI(l922,129;
1924,124)and followedbyallsubsequentauthors. It isnotknownwhatspecimens
ColFFAITusedfor theredescription.SeealsocommentssectionunderQ.sublimbatus.
Quedius(Raphilrus) fellmam(ZETTERSTEDT)
Staphylinusfe11maniZETTERSTEDT,1838,62.
Quediusfe11matli:SMETANA,1971,233.
New record. [Rossia]:SouthernKamchatka:“Mutnovski-Vulkan”,26-VII-91,
R.PREDEL(MSCC)1.
Comments. Thisisthesecondrecordofthisnortherncircumpolarspeciesfrom
Kamchatka(seeSMETANA,1976,28).
VlelleiopsismarginiyentrisFAIRMAIRE
Ve11eiopsismarginiventrisFAIRMAIRE,1882,CLXIV;MARKOVITCH,1915,148-150;COIFFAIT,1978,288.
New record. [Turcia]:Anatoliabor.,cca7kmNWKoso,1,700-2,000m,20-
VII-76,W. HEINZ(ASCC)1.
Comments. Very few recordsofthisveryrare,spectacularspeciesareknown.
ItisatpresentknownfromBulgariaand fromAsiaMinor.
Thecollectioncircumstancesofthisspecimenareunfortunatelynotknown,except
thatithasbeentakeninamixedQuercus-Populusforest.Thetwooriginalspecimens
of thisspecieswerefoundonacliffamongclimbingvines(“sur la falaise,clansun
sentieraumilieudesvignes”,FARIMARIE,1882,CLXV)nearVarna,Bulgaria.Isuspect
that thespeciesmayliveinsubterraneangalleriesofananimal,or innestsofsocial
Hymenoptera.
ve11eiopslsmarginiventris(thetypespeciesofthegenus)agreesingeneralhabitus
and particularly in thequitecharacteristicconfigurationof theantennawith the
Nearcticspeciesof MegaquediusCAsEY, 1915. It ispossible that Ve11eiopsisand
Megaquediusin factrepresentonetaxonthatmaybeamemberof theoldTertiary
fauna.Thishastobeconfirmedbyacomprehensivestudyof thesupraspecifictaxa
