Table Of ContentEl、・1t-a,Tok、o,30(1):235-246,June30,2002
Epaniaopaca(Coleoptera,Cerambycidae)anditsNewRelative,
withaBriefNoteontheGenusMo1orchoepanla
TatsuyaNIIsAT0
BioindicatorCo.,Ltd.,Yarai-cho126,Shinjuku-ku,Tokyo,162-0805Japan
Abstract EpanlaopacaFSIHERisredescribedandillustratedbasedonthetype
specimenspreservedintheNatural HistoryMuseum, London,andistransferredtothe
genusMo1olchoepanla.AnewspeciesrelatedtoopacaisdescribedfromQueenslandof
North East Australia. Thecharacteristicsof the genusMole,choepaniaare brieftyde-
scribed,andachecklistofallthemembersofthegenusisprovided.
Introduction
Mo1orchoepanlaPfc isasmallmo1orchinegroupestablished on thebasisofan
Indochinesespecies,EpanlabarbieriPfc,andonlyfivecongeners haveso farbeen
knowntooccurinEasttoSoutheastAsia.Althoughoriginallydescribedasasubgenus
apoasr of pama, Moforc ﾝoepam show close relationshi t Mote,,c(cid:0)s thn pama
inthebasicstructureoftheprothorax.ThegeneraMotel・choepaniaandMo1orchusare
commonintheautapomorphy inthetribe; that is,“pleuralprocesswidelyseparated
and furcasternumofcoxalcavitieslessdevelopedbehind”(NIIsATo,1986, 1992).It is
doubtlessduetotheindependencyofthegenus,sincethehindwingofMo1olchoepanla
lacksthe3Aveinwhichisalsoautapomorphyofthisgenus.AdultsofMo1orchoepa-
nlaarealmostalwaysfoundonthefreshlydeadvinesandtwigsofhostplants,anddo
notassociatewiththeflowersasthoseoftheothermembersoftheMo1orchini.
InthecourseofmystudyofthetribeMo1orchini, Ihavecontinuouslymadere-
searchesonthetypespecimenspreservedin foreignmuseums.Atanopportunityto
visittheNaturalHistoryMuseum,Londoninthewinterof1999,I foundapairofthe
typespecimensofEpaniaopacaFISHERdescribedfromJavainthecollection.Untilthat
time,Ihadknownthespeciesonly fromFISHER'soriginaldescription,andsimply con-
sidereditasomewhataberrantspeciesofEpania.Tomysurprise,theJavanesespecies
inquestionhastheMo1orchoepanla-1ikefacies,andsoonitbecameevident thatEpa-
nlaopacaactuallybelongstoMotel・choepanlabyacarefulcomparativeexamination
madeundermicroscope. Epanlaopacanot only has the forecoxal cavitieswidely
openposteriadbut alsohasno3A veinofthehindwing. Besides, l also founda fe-
malespecimen fromQueenslandwhichisclosely allied toopacapreserved in the
samecabinet.
Inthefollowinglines,IwilltransferEpanlaopacatothegenusMotel・choepanla
236 TatsuvaNIIsATo
and carefully redescribe it on thebasis of the type specimens, newly describe a
Queenslandspeciesrelatedtoopaca,andthenbriefly discussonthegenus Mo1or-
choepanlaandlistupallthenominatespeciesofthegenus.
Abbreviations. Thefollowingabbreviationsareusedinthedescription:HW-
maximumwidthofheadacrosseyes,FL-lengthof frons,FB-basal widthof frons,
FA-apicalwidthof frons,PA-apicalwidthofpronotum,PB-basalwidthofprono-
tum, PW -maximum width of pronotum, PL-length of pronotum, EW-humeral
widthofelytra,EL-lengthofelytra.
GenusMolorchoepamaPfc,1949
ipama(Mote,c/1oepania)Pfc,1949, Echange,(65),p 9; pespecies: ama a,・bier!PIc, 1949. -
GREsslTT&RoNDoN,1970, Pacif. Ins.Mon.,24,p it2. - NAKANE,1976, Ins. &Nat.,Tokyo,
11(6),p3.
Mole,choepania:HAYAsH1,1974,Ent.Rev.Japan,26,p.15; l984,Coleopt.JapanCol.,Osaka,4,p 55.
- KUsAMA & TAKAKUwA, 1984, Longic.-Beet1.JapanCol., p 282. - NIIsATo, 1986, Ins. &
Nat.,Tokyo,21(12),p9;1992,I11ustr.GuideIdentific.Longic.Beetl.Japan,p 490.
Mo1o1chussubg. Kobaneus HAYAsH1, l958, Ent. Rev.Japan, 9, pp 46-47; type species: Motel'chus
(Line'nlus)'川・oguchiiHAYAsHl,1955.
Thisgenusischaracterizedbyacombinationofthefollowingfeatures:
Bodyrathershortandbroa ratherthickinprofilewithrelativelyshortantennae
intheMo1orchini.Colouralmostalwaysuniformly browntoblack,dull ingeneral,
withoutanymaculationonelytra,thoughsometimeswithpalebandsonventrites1-2.
Hairsandpubescencedense,particularlyonelytra.Headlargeandvoluminous,well
convex,withraised frons,lower lobesofeyessmallandwidelyseparatedfromupper
ones,genaevery deep.Antennae11-segmente slightly longer ( )or fairlyshorter
( )thanbody,notsoslendertowardsapicalsegments,denselyclothedwithminutepu-
bescenceonsixapicalsegments.Pronotumlargeanddistinctlytransverse,simplyar-
cuateatsides,moderatelyconstrictednearapexandbase,notborderedalongapical
margin,withdiscmoreor lessuneventhoughwithoutanycallosities,asperateonsur-
face.Scutellumsmall, triangular.Elytrastronglyreduced,notattainingtothebaseof
abdominal tergite,broadlyroundedatapices,withdiscasperateorgranulateonsur-
face,usually denselyhairednear base.Hindwingsratherbroad, lesselongate,with
simplebutdistinctvein tA3十2A,andwithoutvein3A.Prosternumwith forecoxal
cavitiesdistinctlyangulateexternally,widelyopenposteriad(furcasternumlessdevel-
opedposteriad),presternalprocessdeeplyconcavenearmiddle,andvisibleonlyattri-
angularbaseandcompressedapex.Midcoxalcavitiesbroadlyopentometepisterna.
Abdomensomewhatbroadthoughlessreduced.Legsrelatively longandnotsostout,
withhairyhindtibiae.Medianlobemoderatelyelongate,withapical lobemoderately
convex andbroadly truncateat apex, rather longmedianstruts. Tegmenwithbroad
uni1obedparamere.
Range. Japan (Kyushu, TsushimaIsis., Izu-NiijimaIs.,Ryukyus,etc.), China
(Shaanxi), Taiwan, Indochina (Vietnam, Laos and Thailand), Indonesia (Java and
EpanlaopacaanditsNewRelative 237
Fig. 1. Mole,・choepa,1iaba,・bieri(Pfc,1949), fromnorthernThailand,typespeciesofthegenus. -
,
a,Profile;b,righthindwing;c,prosternum.
Sumatra),NE. Australia(Queensland). I added Sumatratotherangeof thegenus,
sinceanundescribedspeciesofthegenuscollectedfromtheislandisinmycabinet.
Notes. ThoughestablishedasasubgenusofEpanla,Mo1orchoepanlaisconsid-
ered to be an independent genus in recent arrangements (HAYAsHl, 1974, l984;
KUsAMA&TAKAKUwA,1984;NIIsAT0,1986,1992).Thegenushasratheracloserela-
tionshiptothegenusMo1orchusthantoEpanla,becauseof therelativelysmalleyes
and the widely open fore coxal cavities (less posteriorly deve1oped furcasternum)
whichareautapomorphyofthegeneraMo1orc/1usandMo1orchoepaniain thetribe
Mo1orchini.Epanlaisafairlyderivativegenusand formsasister-groupofthegenus
Glaphyt・a; both thegenerahavewelldeveloped furcasternumandexternally recog-
nizedontheposteriorlyclosed forecoxal cavities. Theexternalsimilaritiesbetween
Mo1orchoepanlaandEpanlamaybecausedbyparallelevolutionamongthedifferent
lineagesofthesametribe. It isevident that theindependency of thegenusMo1or-
choepanlainbothmorphologyandecologycanberecognizedbeyondalldoubt.The
membersofgenushaveno3Aveinofthehindwingwhichisaderivativecharacter in
thetribe. Theadultsofthegenusarealmostalwaysactiveonthe freshly deadhost
plantsanddonotvisitflowersincontrasttotheothermembersofthetribe.
Thegenuscontainseightspecies includinganundescribedspecies andwidely
occurs from East Asia viaSoutheast AsiatoNortheast Australia.
238 TatsuvaNIIsATo
Molorchoepanlaopaca(FISHER,1937),combnov
(Figs2&4a)
EpanlaopacaFISHER,1936,Fn.javanica,(79),pp.175-176;typelocality:Koebang-kangkong,Java
Largeandrobustspecies,characterizedbythewhitebandson femaleventrites
1-2.
Female. Colour almostuniformly brown, partly infuscate,slightly dull;head
brown,blackalongsidesandfrontalmarginofclypeus,somewhatinfuscatenearver-
texandtempera,gulalightyellowishbrown;apicesofmandiblesandpalpiyellowish
brown;antennaeunico1oredbrown;pronotumbrown,irregularly infuscateonthedisc,
andalsowithblackbasalmargin;scutellumblackishbrown;ventralsurfaceunicoIored
brown, though lightyellowishbrownonprosternum,whiteinventrites1-2ateach
baseandapical1/3; legsbrown.
Headmoderately voluminousthoughnotsoconvex, rather distinctly narrower
than pronotum, almost entirely covered with close and rather small reticulations,
densely with paleerect hairs, HW/PA 1.05, HW/PW 0.81; fronsslightly narrowed
apica ratherstronglyarcuateatsides,almost attened,slightlydepressednearposte-
riorpartincludingvertex,shallowlyconcavealongmidline,witharcuateapicalmargin
ratherdistinctlyproducedat themiddle,FL/FB0.48,FB/FA1.13;clypeus largeand
transverse,nearly2/5ofbasalwidth,weaklyemarginateatapical margin;genae3/4
thedepthof lowereye-lobes,moderatelyraisedincludingtempera;eyesmoderatein
size,ratherweaklyprominent,withupperlobesseparatedfromeachotherby13/20the
widthofocciput.Antennaefairlylongforamemberofthegenus,0.65timesaslong
asbody,notsoslender,distinctlythickenedapicad,distinctlybroadenedand attened
inapicalsevensegments,moderatelyclothedwithshortbrownishhairsinsegments
1-5, and with dense minute brownpubescence in segments 5-11; scapeweakly
clavate,somewhatdepresse almostequal inlengthtosegment3;segment2strongly
reducedandthickenedatapex;segments3and4thickenedatapex,theformeralittle
morethan3/4thelengthofthelatterwhichisthelongest;segments5-10slightlyde-
creasinginlength;terminalsegmentbluntlypointedatapex.
Pronotumtransverseglobose,almostcompletelyroundedatsides,widestatmid-
dle,stronglyconstrictedat apexandbase,PL/PA t 20, PB/PA 1.03, PL/PW 0.95,
PW/EW0.97,PL/EL0.91;apexweaklyarcuate,verynarrowlybordere a little nar-
rower thanbase;baseverystronglyre?exedandweaklyemarginatenear themiddle,
distinctly bordered; discmoderatelyconvexthough attened inprofile, stronglyde-
pressednear basal collar,withsurfaceirregularlyuneven,providedwithsmall and
closereticulationsinmostpart,denselywithlongerectbrownishhairs,andalsowith
dense recumbent, silvery white pubescence along basal margin. Scutellum small,
roundedatapex,denselypale-pubescent.
Elytradistinctlybroad,stronglyreduced,barelyreachingapical1/5ofmetathorax,
widest atbasal t/4, rathernarrowlyandstraightly dehiscent inapical1/3,distinctly
borderedalongapical thirdofbothexternalandsuturalmargin,EL/EW 1.03; sides
Ep- i1optlcaanditsNewRelative 239
Fig. 2. Mole,choepanlaopaca(Fist-IF_R,1936).combnov.,paratype fromKoebang-kangkong,Java
,
withalmostroundedhumeri,arcuatelyroundedtobasal2/5, thenarcuatelystrongly
convergent toapiceswhicharewidelysubtruncate;discuniformlyconvex,obliquely
raisedfrombasal t/4ofmiddleparttonearapical3/5ofsuture,andratherweaklyso
frommiddleofexternal1/3toapical1/gotsuture,stronglydepressednearsuturebe-
hindscutellum,andobliquelysobetweentworaisedareas,surfacedenselypunctured,
densely clothedwithbrownhairs,andwithpalerecumbenthairsonasubtriangular
partinbasal2/5nearscutellum.
Prosternumalmostflattened,shallowlyfurrowe(i,moderatelywithlongpalehairs;
presternalprocessalmostevanescent,onlyvisibleat theacutelyconvexbase. Meso-
240 TatsuvaNl1sATo
andmetathoracescoarselyandrathercloselypuncture denselyclothedwithlongsj1_
Verywhitehairs. Abdomenmoderatelybroa weakly arcuateatsides,shagreene
clothedwithdensebrownishpubescenceatsidesofventrites1_3.
Le9s,stout,moderatelylong;hind femurmoderatelyclavateinapical 2/3; 1st
hindtarsalsegmentnearlytwiceaslongasthefollowingtwosegmentscombjned.
Bodylength108mm.
Male. Colouruniformlydarkbrown,slightlyreddishonventer,atleastinhind
body,withoutwhitebandsonabdominalsternites.Elytrasimilartothoseof female,
though narrower, with humeri moderately expanded, apices completely rounde
EL/EW0.88;discwithwideobliquedepressionfrombasal t/3nearexternalmarginto
apical1/3ofsuture,puncturedandpubescentasin female,thoughsupplementarily
providedwithdensesilverywhitepubescenceatatriangularpart inbasal4/7near
scutellum.Ventralsurfacealmostasin female,thoughsilverywhitepubescence fairly
denseat thesidesofventrites. Legsslightlylongerandstouterthanin female.Body
length7mmaccordingtoFISHERs' originaldescription(headandprothoraxaremiss_
ingintheholotypeexamined).Malegenitalianotexamined,becauseofthepoorcon_
clitionofthespecimen.
Specimensexamlrled. 1(3(holotype),“S.C.DRESCHER”“Koebang-kangko_
engZuid-Banjoemas,Java,25.Mr'17.1.1932'(whitelabel)/“Brit.Mus.1937_662.”
(whitelabel)/“dot:19_Epaniaopaca1933/36FISHER”(pinklabelwithblackmargin);
1 (paratype),almostofthesamedataasthepreceding,butthecollectingdatejs“19.
1.1932”.Theholotypespecimenexaminedlackstheheadandprothoraxincludingap_
Penda9es,excepttheleft forelegwhichispastedonthemountingcard.Theparatype
femaleisinafaircondition,thoughtheapicalsixsegmentsofthele量antennaandthe
leftlegaremissing. Imadetheaboveredescriptionmainlyonthebasisofthe female
paratype forthereasonoftheconditionofthetypespecimens.Thesetypespecimens
arepreservedintheNaturalHistoryMuseum,London.
fstrzbtffzon. Java,Indonesia.
Notes. Thisisaveryuniquespeciesinhavingrobusthabituswith large fore
body, theapicallythickenedantennaeand thewhitishbasalsegmentsof the female
ventrites.Thesecharacteristicsarenotobservedintheothercongenersofthegenus.
However, thisspeciesbelongstothegenusMo1orchoepanlabeyondalldoubt,since
suchautapomorphyofthegenusasthesmalleyes,thelessdevelopedfurcasternumof
theforecoxalcavities,thesimplyarcuatesidesofpronotum,andlackofthe3Avein
ofthehindwingareallpresent.
Mo1orchoepanlaopacacombnov.islittleknownspeciessinceonlyapairofthe
typespecimenswereexaminedatthepresenttime.Noadditionalspecimenhasnotyet
beenobtainedfromJavainaboutpastseventyyears.
Epa,fiaopacaanditsNewRelative 241
Molorchoepanlaalbiventrissp nov
(Figs 3&4c)
Asmallspecieswithlarge forebody,characterizedbyentirelyyellowishwhite
abdominalventrites1-2.
Female. Colour dark yellowishbrown todark reddishbrown,weaklyshiny;
headdarkreddishbrown,infuscatealonganteriormarginoffronsandgenae,andalso
longitudinallyreddishnearmidlineofgula;apicesofmandiblesandpalpiyellowish
brown,eyesblack;antennaeyellowishbrownatleastinscapeandsegment2;prono-
tumdarkreddishbrowntoreddishbrown,withblackasperations;scutellumyellowish
brown;elytrayellowishbrown,withblackishasperation;prosternumreddishbrown,
slightlyyellowishnearmiddle;meso-andmetathoracesdark yellowishbrown, more
yellowishatmesosterna1process,paleronmedianproducedpartofmetasternum;ab-
domenyellowishwhiteinventrites1-2,andblackishbrownonventrites3-5though
largelyreddishnearmiddle;legsunico1orouslyyellowishbrown.
Headmoderatelylargeandvoluminous,whollywellconvex,ratherdistinctlynar-
rower than pronotum, closely covered with shallow and rather small reticulations
above,almostsmoothatsidesand venter,moderatelycoveredwithshortpalehairs,
HW/PA0.80,HW/PW0.71; fronssubquadrate,lessnarrowedapicad,gentlyarcuateat
sides,moderatelyandevenlyconvex,withsimplyarcuateapicalmargin,FL/FB0.68,
FB/FA 1.04;clypeusmoderate, transverse,nearly 1/4 thebasal width,gently emar-
ginateatanteriormargin;genae3/4thedepthoflowereye-lobes,weaklyraised;eyes
relativelylarge,moderatelyprominent,withupperlobesseparatedfromeachotherby
2/3 thewidthofocciput; vertexandocciput distinctly raised. Antennaewithscape
weaklydilatedapica depressed(mostsegmentsaremissingintheholotype).
pronotumsomewhatglobose,almostaslongaswide,widestjustbeforemiddle,
distinctlyarcuateatsides,morestronglycontractedtobasethantoapex,PL/PA1.16,
PB/PA0.80,pL/pw1.03,pw/Ew1.00,pL/EL1.00;apexalmost transverselytrun-
cate,notbordere distinctlywiderthanbase;basenarrowlybordere weaklyarcuate
andslightly emarginatenearmiddle; discmoderately convex, inprofile attened in
apical1/4,thenstraightlydeclinedtobasal t/10,unevenabove,providedwithanindis-
tinctswellingatmiddleofapical3/10andapairofswellingsatsidesofbasal1/5;sur-
facemoderatelyprovidedwithsmallasperation,clothedwithpalehairs,thehairspar-
ticularlydenseatsides,andalsowithdensesilverywhitepubescenceatwholesides.
Scutellum relatively large,moderatelynarrowed toapexwhich is rounded, densely
clothedwithsilverywhitepubescence.
Elytramoderatelybroad,stronglyreduce almostreachingtheleve1ofhindcoxal
cavities,notsostronglynarrowedapicad,widestatbasal t/3,straightlydehiscent in
apical 1/2, indistinctlyborderedorsimpleat least nearapices,EL/EW 1.00; sides
ratherstronglyexpandedathumeri,weaklyarcuatetojustbeforemiddlethenweakly
andarcuatelyconvergenttoapiceswhicharenarrowlyrounded; discuniformlycon-
vex,moderatelyraisednearsuturebehindscutellum,alongobliquepartsfromhumeri
242 TatsuyaNllsAT0
Fi9. 3. Mole'choepanlaalb、i'ent''isspnov.,holotype fromKuranda,Queensland,NE.Australja
,
tobasalhalfof themiddleofdisc,andatapical1/5;surfaceshagreene partlypro_
Videdwithasperationmostlyontheraisedparts,denselyclothedwithbrownhairs,the
hairsparticularlydenseonbasal2/5nearsutureandatsidesofbasal3/5.
P「oSternumslightlyraisedtowardsapex,almostsmooth,sparselyclothedwjth
PaleIon9hai「S;Presternalprocessalmostevanescent,onlyvisibleattheacuteconvex
base.MeSo-andmetathoracesshagreened,partlywithcoarsepuncturesonmesoster_
num,afewSmallonesonposteriorpartofmetasternum,moderatelyclothedwithpale
longhairs,andalsodenselywithsilverywhitepubescenceatsidesofmesothoraxand
atPosteriorPartofmetepisternum.Abdomenratherlong,weaklyarcuateatsides,gent_
1yema「9inateatapexofanalventrite,aml ostsmooth,rathersparselyclothedwjthpale
longhairs.
Legsmoderatein length,stronglycompressedin femoraandtibiae;hjnd femur
EpaniaopacaanditsNewRelative 243
c
lg~ -
lc,3S/3
t j'
f- 1
0,1_3,一 ｫ Brit.Mui・
l' ;Z-,-:- r一・一一 ・ l9 2。
b
.
Fig. 4. Photographsof thetypespecimensofMole,-c1loepa川aspp. - a, M opaca(FISHER,1936)
comb nov., paratype from Koebang-kangkong,Java;b,ditto,labels;c, M albi、,e,It・,Issp nov.
,
holotype fromKuranda,Queensland,NE.Australia.
,
slightlyclavateinapical5/12;hindtibiaprovidedwithdentateasperation inapical
3/11alongexternalmargin.
Bodylength5.8mm.
Type specimen. Holotype “Kuranda, N. Queensland, G. E. Bryant. P. D.
,
Dori 1911”(whitelabelwithredtransverselineatmiddle)/“G.BryantCol1. l919-
147”(whitelabel)/“Br.Mus.(reddishpinklabel)”/“?Epanlasp”“Id.ByH.J.Carter”
(whitelabel).Thespecimenexaminedisinafairlypoorcondition.Boththeantennae
excepttheleftscape,andtherightscapeandpedicel, theleftclawandtarsalsegment
3,thelefthindleg, therighthindtarsi,andthele量hindwingaremissing.Theholo-
typeispreservedintheNaturalHistoryMuseum,London.
Distrtbution. Queenslan NE. Australia.
244 TatsuyaNIIsAT0
Notes. Mo1orchoepanlaalblventrissp nodoubt belongsto thesamelin-
nov
eageasM.opacacomb nov.,andshareswiththelattersuchuniquecharactersasthe
largeanddistinctlyglobularpronotum, thewideandstrongly reducedelytra, and
whitishventrites1-2.Thethickenedapicalsegmentsoftheantennaemayalsobecom-
moninthetworelatives,butIwasunabletorecognizethischaractersinceboththean-
tennaearemissingintheholotypeofthenewspecies.Mo1orchoepanlaalbiventrissp
novandM.opacacombnovmaycomposeaspecies-groupinthegenus.
Thisnewspecies isthefirst representativeofMo1orchoepanlaknown fromthe
AustralianSubcontinent,andalsothesouthernmost inhabitantof thegenus. 0nly a
singlefemalespecimen inaverypoorconditionwasunfortunatelyexamined.How-
ever,Idescribeditinviewofitspeculiarityandaffinityofthespecies.Thenewname
isderivedfromthewhitebaseoftheventritesofthespecies.
ChecklistoftheGenusMolorchoepamaPfc
Mofore(cid:0)epam arOfen(PIc,1949)
Epanla(Mo1o1choepanla)ba・,hie,・iPIc,1949,Echange,(65),p9;typearea:Cochin-China. - GREss1TT
&RoNDoN,1970,Pacif Ins.Mon.,24,p it2,fig21c;localityrecord:C.Laos.
Mole・,choepanlaba,-bie,-i:HAYAsHI,1974,Ent.Rev.Japan,26,p.15.
zsfn utzon. S.Vietnam,LaosandThailand(newrecord).
ma, Moforc(cid:0)epam cooma (Pfc 1940
EpanlacoomaniPfc,1940,Echange,(56),p4;typearea:Tonkin.
Epania(Mole,choepanla)cooman1:GREsslTT&RoNDoN,1970,Pacif.Ins.Mon.,24,p it2,fig 21d;10-
calityrecord:PhonTiouandVientian,Laos.
Mo1orchoepanlacoo,nan1:HAYAsHI,1974,Ent.Rev.Japan,26,p. l6.
fstn ufzon. N. VietnamandLaos.
Mo1orchoepanlasimplec)a(MATSUSHITA,1933)
Mo1orchusslmplexusMATSUSHITA,1933,J.Fac.Agric.Hokkaidoimp.Univ.,34,p 228,pl 3, figs.1a,b
typelocality:Hozan,Formosa. - GREssITT,1951,Longicomia,2,p.173.
Mole,chus(Slnolus)simplexus:MIToNo,l940,p 62.
Mole'chus(s.str)simpiexus:HUA, l982,CheckListLongic.Beetl.ChinaColeopt.Cerambyc.,p42.
Mole'choepaniasimplexa:NAKAMURA,S.,eta1.,1992,Check-listLongic.-beetl.Taiwan,p31.
Distribution. Taiwan.
5aI Mofore(cid:0)epam mzzogtlc(cid:0)z (HAYAsH, 195
Mole'chus(Linomitls)mizoguchiiHAYAsHl,1955,Col.I1lustr.Ins.Japan,1,pp51,54,pi t9, fig204;type
locality:SataCape,KagoshimaProf.
Mole'chus(Kobaneus)'nlzoguchit:HAYAsHI,1958,Ent.Rev.Japan,9,pp 46-47. - 0HBAYAsH1, l963,
