Table Of Content^Ssou
American Fern Journal 92(l):l-9 (2002)
Rl
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May
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Milton Tryon,
Rolla
Obituary:
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(1916-2001)
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Gastony
Gerald
J.
47405-3700
Bloomington, IN
Indiana University,
Department of Biology,
Barrington
David
S.
VT
05404-0086
Vermont, Burlington,
University of
Department of Botany,
Con ant
David
S.
VT
05851
Lyndon Lyndonville,
State College,
Department of Natural Sciences,
and one
member American Fern Society since 1932 of
of the
Rolla Tryon, a
was born on
most eminent students of pteridophytes,
twentieth century's
the
American
His professor of history
father, a
August 1916 Chicago, Illinois.
in
26,
summer
Chicago, maintained a cottage in
University of
and education the
at
home
Chicago. RoUa's fascination
addition his in
to
Indiana in
Chesterton,
boyhood from Ches-
developed during forays that
and
with fern allies
ferns
Hodge Mexico December, 1972 and was
was taken by Dr. Walter H. in City in
The photograph
Documentation.
Hunt Botanical
made by the Institute for
available
AMERICAN
FERN VOLUME NUMBER
2 JOURNAL:
92
1
(2002J
terton cottage into sand dune habitats along Lake Michigan in the northwest
At
of Indiana. the age of he piibUshed
18, his first paper, relating his obser-
Osmunda
on
vations Dunes
plants in the Indiana complete
(see bibliography
As
below). was
a boy, Rolla greatly influenced and
by, in turn influenced,
Deam
Charles (author of the 1940 Flora of Indiana], advising Deam about fern
species he had found dunes When Deam
in the area. a doubting appeared
at
the cottage door one day asking meet and shown
Rolla be
to to these ferns in
he was
surprised was
situ, to learn that Rolla not the adult of the family but
mere boy Thus
a of 14. began a productive friendship documented
in Rolla's
Deam
correspondence
with from May, 1935 penny
to January, 1953. All the
postcards and he Deam
received from have been
letters maintained
carefully
—
now
one
in of Rolla's files, archived at Indiana University fascinating reading.
boyhood
him
Rolla's insatiable appetite for ferns got into a bit of trouble
at
home, when
however,
his father learned that he had charged Bower's
three
volumes on The
Ferns account
to his Brentano's bookstore
at in Chicago.
Rolla built a solid academic superstructure on the foundation of these boy-
Among
hood
experiences. his academic accomplishments were an A.A. degree
and
1935
in a B.S. degree in 1937, both from the University of Chicago, and
Ph.M. 1938 from
a in the University of Wisconsin. In 1940 he earned an M.S.
and
in 1941 a Ph.D., both from Harvard
University. During his days as a Har-
vard he
student, contracted malaria South
in Carolina while
collecting plants
M
Warfare
Massachusetts
Ph.D. botany with
in another, time
this in chemistry, so that he could earn
a
Dartmouth
Wis
Minnesota
While
in 1945.
Wis
in 1945 initiated not only happy and enduring
a domestic
partnership but
also
a research synergism whose
productivity has nourished
pteridologists through-
out the world.
became
In 1947 Rolla Associate Professor Botany Washington
in Univer-
at
and
sity, St. Louis Assistant Curator of the herbarium of the Missouri Botanical
Garden, positions he held 1957. During appointment, he and
to this Alice were
the original organizers of the Missouri Botanical Garden's annual
Systematics
Symposium, whose
48th meeting was held 12-13
October, 2001. This highly
successful annual meeting has received continuous support from
the National
Science Foundation from second
year
its (1954) to the present (with the lapse
From
of a single 1946 1957
year). Rolla served and
to as curator librarian of
American
the Fern and
Society's library herbarium, responding members'
to
requests for loans of materials. That herbarium and was
library subsequently
entrusted Warren H. Wagner
to at the University of Michigan. Following
a year
Research
as Associate
the University
at of California, Berkeley, Rolla went
to
Gray Herbarium
the Harvard
of University as Associate Curator and
Curator
of Ferns in 1958 and became Curator of the Gray Herbarium
in 1967. Rolla
ROLLA MILTON TRYON,
3
JR.
and Alice traveled the world extensively, attending international meetings,
conducting field work, studying specimens at major herbaria in the Americas,
Europe, and Africa, and conducting field courses on the ferns. In addition to
many
other services to professional societies, Rolla served for years as Asso-
ciate Editor of Rhodora and the American Fern Journal, as Associate Editor of
and
Brittonia (1961-1964), as Editor-in-Chief of fli7odora (1977-1982), as Pres-
New
ident of the England Botanical Club and the American Fern Society.
A
framed photograph of his revered mentor, Charles A. Weatherby (see
American Fern Journal 40[ll for a remarkable series of papers honoring this
unusually respected and beloved botanist), was always prominently displayed
own
on Rolla's desk Harvard, undoubtedly inspiring his welcoming, patient,
at
who
and supportive response to all entered his office seeking counsel. In 1970
New
Rolla an annual England Fern Conference Harvard Forest. For
initiated at
which
20 years this provided a stimulating intellectual setting in students of
became
fern biology discussed and developed their ideas. In 1972 he Professor
and
of Biology Harvard University, holding both the Curatorship Professor-
at
He
ship until his retirement in 1987. remained at Harvard as Professor Emer-
when moved
itus from 1987 to 1989 he to the University of South Florida in
Tampa him
as Adjunct Professor, bringing with his extensive library of fern
and biogeographic To mark the retirements of Alice and Rolla Tryon
literature.
from Harvard, of 13 papers plus introduction was published in
a festschrift
honor Annals Missouri Botanical Garden 225-339.
their in the of the (vol. 77:
1990).
At the University of South Florida, Rolla and Alice helped found the Insti-
endowed
Systematic Botany and the Tryon Lecture Series that brings
tute for
known
each
several internationally botanists to the university year. In their
on Tampa campus, he and Alice continued
research-active the their pter-
office
idological work, as his following bibliography indicates.
exceeds 100 and includes breadth
Rolla Tryon's publication titles a great
list
of topics. Papers ranged from articles on pteridophytes for the 1943 Encyclo-
pedia Britannica to a glossary of terms relating to the fern leaf, discussions of
the history of pteridology and fern classification, a remembrance of his grad-
mentor and counselor Charles A. Weatherby, discussions of the formali-
uate
ties of fern nomenclature, and many book reviews. His monographs and re-
visions focused mostly on ferns but also included angiosperms Convolvulus
among were
and Elymus. Signal these his revisions oi Pteridium, Doryopteris,
American Notholaena, and Cyatheaceae.
group, the
the Selaginella rupestris
on biogeography began with Doryopteris in 1944, matured in
His papers fern
and continued
his exposition of geographic speciation in Selaginella in 1971,
phytogeography North
and 1999 discussion of the of eastern
to his Alice's
American (honoring Ching Ren-Chang). Floristic and taxonomic notes on
ferns
and
growth forms hybrids
ranged from simple observations of to eluci-
ferns
dations of complex taxonomic and nomenclatural issues. For his 1955 publi-
Washington
cation on the taxonomy of cycads (coauthored with students in his
University he was awarded the 1956 Robert Montgomery award of the
class)
Garden distinguished achievement in the world of palms
Fairchild Tropical for
AMERICAN FERN JOURNAL: VOLUME NUMBER
4
92
1 (2002)
and cycads. At the time of his death, he had a book review in press in Rhodora
and a paper in press in Bradea (coauthored with his former student Paulo
Windisch).
numerous
In addition to his papers on ferns and other topics, Rolla no-
is
Among
table for his books. these are two editions of his Ferns and Fern Allies
Wisconsin 1953) and Ferns and Fern Minnesota
of (1940, Allies of (1954,
He
renowned knowledge
1980). is for his of the ferns of Peru, expressed
first
in his 1964 Ferns of Peru (250 pages in the Contributions from the Gray Her-
barium Harvard This treatment was updated and completed
of University). in
six subsequent parts entitled Pteridophyta of Peru between 1989 and 1994,
mostly coauthored with Robert Stolze but with several portions contributed
by other pteridological His monumental 1982 book Ferns and
specialists. Al-
lied Plants with Special Reference to Tropical America, coauthored with Alice
Tryon, an encyclopedic
is treatment of this subject that continues to stimulate
new
research, as does his treatment of Pteridaceae, with Alice Tryon and Karl
Kramer, volume and
in 1 of Families Genera of Vascular Plants edited by K.
R
Kramer and
Green,
S.
and
Rolla's kindly perceptive mentoring and his outstanding contributions
knowledge
to our of ferns is signaled by having the following four fern taxa
named
in his honor.
Asplenium
1) tryonii CorrelL In describing this species, Donovan Correll
name who
(1961) said '*It is a pleasure to this species for Dr. Tryon, has
always been most
gracious in helping his fellow-workers with their never-
Known
ending problems
in the study of ferns/' only from Chihuahua, Mex-
was
ico, this species further discussed and illustrated in Ferns and Fern
'/
tryonorum The eminent Mexican
Riba.
Journal 90:112-118, 2000)
stated that "this species
is
M
taxonomy
ognizes the close professional relationship between this highly productive
now known
research team. This
tree fern species as Trichipteris tryono-
is
rum
Tryon by
(Riba) R. following transfer Rolla in his 1970 paper on the
its
classification of the Cyatheaceae.
am my
Nephelea name
tryoniana Gastony. pleased
*T to this species for
mentor, M.
Dr. Rolla Tryon, in recognition of his outstanding contribution
the understanding and
to of the systematics evolution of the family Cy-
atheaceae" (Gastony, 1973). Subsequent research by Conant (Conant and
Cooper-Driver, 1980; Conant, 1983) revealed
that this tree fern species a
is
now
reproductively stabilized diploid hybrid species regarded
that as
is
Alsophila tryoniana
(Gas
Tryonella Pichi Sermolli
4)
M
many
author of important papers on who,
supported
ferns, inter alia, the
distinction of the present genus from Doryopteris, though without giving
it
ROLLA MILTON
TRYON,
JR. 5
Dory
opt
Kramer
Among
the doctoral graduate students he trained, Rolla counted the follow-
ing (those with asterisks received their degrees from other institutions): Alice
Ramon
Tryon, *Karl Kramer, * Riba, Gerald Gastony, Lawrence Palkovic,
F.
David Barrington, David Conant, Paulo Windisch, James Hickey, *Robbin
*R.
He
Moran, Sonia Sultan, and Galvin Sperling. also mentored Robert Stolze in
Museum.
taxonomic Cnemidaria Always
his revision of the Field available
at
to his students, he modeled his supportive and insightful mentoring on his
own
experiences with graduate mentor, Charles Weatherby. For he has
his this
earned our love as well as our respect. His impact on his students, and their
and
students, their students incalculable.
is
1978 M. was honorary membership Amer-
In Rolla Tryon, elected to in the
Jr.
who
ican Fern Society, a special category of membership for persons have
made
outstanding contributions to the study of ferns. In 1984 he received a
Award
Merit from the Botanical Society of America recognition of distin-
**In
guished achievement in and contributions to the advancement of botanical
science. Pre-eminently knowledgeable in matters of taxonomy and nomencla-
ture, this foremost pteridologist a perceptive student of phytogeography and
is
of the evolutionary impact of the selective process during plant migration."
was
he
In addition to Rolla's botanical activities also highly skilled in run-
many
Knox He
ning a family farm in County, Indiana for years. visited the
farm, overseeing management, a few times each year. The extensive records
its
he managing and man-
kept in crops livestock illustrate his practical ability in
aging business as well as scientific data.
On
August 20, 2001, six days before his eighty-fifth birthday, Rolla Milton
work
Tryon, us continue our with the pteridophytes of the world,
to
left
Jr.,
We
and delight in them, without him. do this fortified by his writings, the
to
He
echoes encouraging words, and his everlasting example. was the
of his
beloved husband of Alice Faber Tryon, the benefactor of countless students of
many who knew him
never an
pteridophytes, including personally, inspiration
many and
and counselor collaborators coauthors, the advisor of doctoral
to
and and
students, the teacher of innimierable undergraduates, our dear friend
He
He
mentor. be deeply missed. already
will
is.
Literature Cited
A
CoNANT, D. 1983. revision of the genus Alsophila (Cyatheaceae) in the Americas. Arnold
S.
J.
333-382.
Arbor. 64:
CoNANT, and Coofer-Driver. 1980. Autogamous allohomoploidy in Alsophila and Ne-
D. G.
S.
new homoploid homosporous
pbelea (Cyatheaceae): a hypothesis for speciation in ferns.
Amer. 1269-1288.
Bot. 67:
J.
Two Texas-Chihuahuan Wrightia 200-203.
CuRRELL, D. 1961. ferns. 2:
S.
A
Gastony, G. 1973. revision of the fern genus Nephelea. Contr. Gray Herb. 203: 81-148.
J.
Knobloch, W. and Correll. 1962. Ferns and fern allies of Chihuahua, Mexico. Texas
D. S.
I.
Research Foundation, Renner, Texas.
Webbia
Fragmenta pteridologiae— 1-16.
PiCHi Sermolli, R. E. G. 1974. V. 29:
AMERICAN FERN VOLUME NUMBER
JOURNAL:
6 92
1 (2002)
New
RiBA, R. 1967. taxa in the genus Alsophila. Rhodora 69: 65-68.
Bibliography of Rolla M. Tryon, (1916-2001)
Jr.
Some
Tryon, R. M., 1934. observations on Osmundas. Amer. Fern 24: 119.
Jr.
J.
Tryon, R. M., 1936. Ferns of the dune region of Indiana. Amer. MidL Naturalist 425-429.
17:
Jr.
Tryon, R. M., 1936. Botrycbium dissecturn mid form a obliquum. Amer. Fern 26: 26-30.
Jr.
J.
Tryon, R. M., 1938. The phenomenon of forking in ferns. Amer. Fern 28: 81-90.
Jr.
J.
Tryon, R. M., 1938. Recent additions to the flora of Indiana. Proc. Indiana Acad. Sci. 47: 76-
Jr.
77.
Tryon, R. M., 1939. Notes on the ferns of Wisconsin. Amer. Fern 29. 1-9.
Jr.
J.
Tryon, R. M., Jr. 1939. The varieties of Convolvulus spitharnaeus and of C. sepium. Rhodora 41:
415-423.
Tryon, R. M., 1940. Notes on some Indiana plants. Proc. Indiana Acad. Sci. 49: 89-90.
Jr.
Tryon, R. M., N. C. Fassett, D. W. Dunlop, and M. E. Diemer. 1940. The ferns and fern allies
Jr.,
of Wisconsin. Botany Dept., Univ. of Wisconsin, Madison.
An Osmunda
Tryon,
R. M., 1940. hybrid. Amer. Fern 65-66.
30:
Jr.
J.
A
Tryon,
R. M.. 1941. revision of the genus Pteridlum. Rhodora 43: 1-31, 37-67. Reprinted as
Jr.
Contr. Gray Herb. 134: 1-70.
Tryon, R. M., Jr. 1942. Review of Roland, A. E., The ferns of Nova Scotio. Amer. Fern 32: 73-
J.
74.
A
Tryon, new
R. M., 1942. Dryopteris hybrid. Amer. Fern 81-85.
32:
Jr.
J.
A
Tryon,
R. M., 1942. revision of the genus Doryopteris. Contr. Gray Herb. 143: 1-80.
Jr.
Tryon, R. M., Jr. 1942. Review of Brown, C. A. and D. S. Correll. The ferns and fern allies of
Louisiana. Rhodora 484-485.
44:
Tryon,
R. M., Jr. 1943. Several articles on Pteridophyta for Encyclopedia Brittanica.
Tryon, R. M., Jr. 1943-1947. Abstracts of the American Fern Journal for Biological Abstracts.
Tryon, Dynamic
R. M., 1944. phytogeography of Doryopteris. Amer. 470-473.
Jr. Dot. 31:
J.
Tryon, R. M., and W. Moore. 1946. Notes on aquatic and prairie vegetation in southwestern
Jr.
J.
Minnesota. Rhodora 113-116.
48:
A
Tryon, and W. Moore.
R. M., Jr. 1946. preliminary checklist of the flowering plants, ferns
J.
and
fern allies of Minnesota. Univ. of Minnesota, Minneapolis.
A
Tryon, new
R. M., 1946. Doryopteris hybrid. Amer. Fern 48-49.
Jr. 36:
J.
A
Moore, W. and M. Tryon, new
R. 1946. record of Isoetes melanopoda. Amer. Fern
Jr. 36:
J.
J.
89-91.
A
Butters, F. K. and R. M. Tryon, Jr. 1948. fertile mutant of a Woodsia hybrid. Amer. Fern 35:
J.
132.
Tryon, R. M., 1948. Some Woodsias from the north shore of Lake Superior. Amer. Fern
Jr. 38:
J.
159-170.
A
Booher, L. E. and R. M. Tryon, 1948. study of Elynms in Minnesota. Rhodora 50: 80-91.
Jr.
—
Tryon, R. M., 1950. Charles Alfred Weatherby teacher and counselor. Amer. Fern 9-10.
40:
Jr.
J.
A
Tryon, new
R. M., Jr. 1950. erect species of tlie Selaginella rupestris group. Amer. Fern 40:
J.
69-74.
Tryon, R. M., Jr. 1951. Ferns of the Missouri Ozark region. Missouri Bot. Card. Bull. 39: 136-138.
Tryon, R. M., Jr. 1951. Review of Manton, I., Problems of cytology and evolution in the Pterido-
phyta, Ecology 769-770.
32:
A
Tryon,
R. M., Jr. 1952. sketch of the history of fern classification. Ann. Missouri. Bot. Gard. 39:
255-262.
Tryon, R. M., Jr., N. C. Fassett, D. W. Dunlop and M. E. Dfemer. 1953. The ferns and fern allies
Wisconsin, Ed. Univ. Wisconsin Madison.
of Press,
2.
Tryon, R. M., Jr. 1954. The ferns and fern allies of Minnesota. Univ. Minnesota Press, Minneap-
olis.
Tryon, Barbour,
R. M., Jr, E. E. Davis, H. Kidd, R. Long, C. Marvin, B. Mikula, and R. Mohlen-
brock. 1955. Ancient seed plants: the cycads. Missouri. Bot. Gard. Bull. 43: 65-80.
ROLLA MILTON
TRYON,
JR. 7
Tryon,
R. M., Jr. 1955. Selaginella rupestris and its allies. Ann. Missouri. Bot. Card. 42: 1-99.
A
Kramer, K. U. and R. M. Tryon, 1955. new species of Doryopteris from Surinam. Ann.
Jr.
Missouri. Bot. Card. 42: 213-214.
A
Tryon,
R. M., Jr. 1956. revision of the American species of Notlwlaena. Contr. Gray Herb. 179.
4-106.
Tryon, R. M., Jr. 1957. Adlanturn in Peru: new species and combinations. Anier. Fern 47: 139-
J.
143.
KoBUSKi, C. E., C. V. Morton, M. Ovvnbey, and R. M. Tryon. 1958. Report of the committee
for
recommendations on
desirable procedures in herbarium practice and ethics. Brittonia 10:
93-95.
Tryon, R. M., Jr. and A. F. Tryon. 1959. Observations on cultivated ferns: the hardy species of
tree ferns (Dicksonia and Cyatheaceae). Amer. Fern. 49: 129-142. Reprinted in Lasca Leaves
J.
26-33. 1960.
10:
Tryon, The
R. M., 1960. ecology of Peruvian Amer. Fern 46-55.
ferns. 50:
Jr.
J.
A
Tryon, some
R. 1960. glossary of terms relating to the fern leaf. Taxon 9: 104-109. Reprinted
in Russian translation In Bot. Jour. Akad. Nauk, U.S.S.R. 46: 736-739. 1961.
A
Tryon, Review Munz,
R. M., 1960. of Pteridophyia in A., CaUfornia Amer. Fern
Jr. P. flora.
J.
157-159.
50:
A
Tryon,
R. M., 1960. review of the genus Dennstaedtia in America. Contr, Gray Herb. 187:
Jr.
23-52.
New
Tryon, R. M., 1960. species of ferns from Central and South America. Rhodora 1-10.
62:
Jr.
Tryon, R. M., 1961. Taxonomic fern notes. Rhodora 63: 7-88.
Jr.
I.
Tryon, R. M., 1962. The fern genus Doryopteris in Santa Catarina and Rio Grande do Sul,
Jr.
Sellowia 51-59.
Brazil. 14:
Tryon, R. M., 1962, Review of Wherry, E. T., The Fern Guide. Amer. Fern 52: 89-91.
Jr.
J.
A
Tryon, R. M., 1962. note on Nepbrolepis cordifolia cv. Daffii. Amer. Fern 52: 153-155.
Jr.
J.
Tryon, R. M., 1962. Taxonomic Fern Notes. PityTogromma (including Trismerio) and Ana-
Jr. II.
gramma. Contr. Gray Herb. 189. 52—76.
Tryon, R. 1962. Taxonomic fern notes. Contr. Gray Herb. 191: 91-107.
III.
Tryon, R. M., 1962. Review of Knobloch, W. and D. S. Correll, Ferns and Fern Allies of
Jr. I.
Chihuahua. Rhodora 347-348.
64:
A
Tryon, R. 1962. commentary on superfluous names. Taxon 11: 116-120.
Tryon, Nomenclatural Taxon 28-285.
R. 1963. proposals. 12:
Mem.
Tryon, R. M. 1964. Evolution in the leaf of living ferns. Torrey Bot. Club 21: 73-85.
Tryon, R. 1964. The ferns of Peru. Polypodiaceae (Dennstaedtieae to Oleandreae]. Contr. Gray
Herb. 194: 1-253.
Tryon, R. 1964. Taxonomic Fern Notes IV. Rhodora 66: 110-117.
Pichi-Sermolli, R. E. G., F. Ballard, R. E. Holttum, H. Ito, F. M. Jarrett, A, C. Jermy, E. A. C.
L. E. Schelpe, M.-L. Tardieu-Blot, and R. M. Tryon. 1965. Index Filicum Supplementum
Quartum pro Annis 1934-1960. Internat. Bureau for Plant Taxonomy and Nomenclature,
Utrecht, Netherlands.
Tryon, R. 1965. Paraphyses in the ferns. Taxon 14: 213-218.
A
Tryon, R. and D. M. Britton. 1966. study of variation in the cytotypes of Dryopteris spinulosa.
Rhodora 59-92.
68;
New
Tryon, R. 1967. Taxonomic fern notes V. combinations in Peruvian species of Thelypteris.
Rhodora
5-8.
69:
Tryon, R. 1967. Review of Hara. H., The flora of eastern Himalaya, Rhodora 69; 456-457.
A
compounds some
BojiM, B. A. and R. M. Tryon. 1967. Phenolic in ferns. survey of ferns for
I.
cinnamic acid and benzoic acid derivatives. Canad. Bot. 45: 585-593.
J.
Tryon, R. M. and A. F Tryon. 1968. Edith Scamman (1882-1967J. Amer. Fern 58: 1-4.
J.
Tryon, R. 1968. Nomenclatural proposals. Taxon 17: 588-590.
Ewan A
Tryon, R. 1969. Pteridology, Pp. 97-102 in (ed.) short histor}' of botany in the United
J.
New
Hafner Publishing Co., York.
States.
Tryon, R. 1969. Taxonomic problems in the geography of North American ferns. BioScience 19:
790-795.
VOLUME NUMBER
AMERICAN FERN JOURNAL:
8 92 (2002)
1
Manual
Tryon, R. 1970. Review of Correll, D. S. and M. C. Johnston., of the vascular plants of
Rhodora 533-535.
Texas. 72:
Tryon, R. 1970. The classification of the Cyatheaceae. Contr. Gray Herb. 200: 3-53.
Tryon, R. 1970. Development and evolution of fern floras of oceanic islands. Biotropica 2: 76-84.
Reprinted 1971 as Pp. 54-62 in W. L. Stern (ed.) Adaptive aspects of insular evolution.
Washington
State Univ. Press.
Tryon, R. 1971. The American tree ferns allied to Spbaeropteris horrida. Rliodora 73; 1-19.
Tryon. R. 1971. Ferns of the Andes and Amazon. Morris Arbor. Bull. 22: 7-13.
The 89-
Tryon, R. 1971. process of evolutionary migration in species of Selaginella. Brittonia 23:
100.
Tryon, R. 1972. Endemic areas and geographic speciation in tropical American ferns. Biotropica
121-131.
4:
—
New
Tryon, R. 1972. Taxonomic fern notes, VI species of American Cyatheaceae. Rhodora 74:
441-450.
Tryon, R. M., and A. Tryon. 1973. Geography, spores and evolutionary relations in the
F.
Jr.
cheilanthoid ferns. Bot. Linn. Soc. 67: Suppl. 1; 145-153.
J.
65-
Tryon, A. and R. Tryon. 1973. Thelypteris in northeastern North America. Amer. Fern 63:
J.
76.
Tryon, R., B. Voeller, A. Tryon, and R. Riba. 1973. Fern biology in Mexico (a class field program).
BioScience 28-33.
23:
Tryon, Widen, A. Huhtikangas, and M. Lounasmaa. 1973. Phloroglucinol derivatives in
R.,
C.-J.
Dryopteris parallelogramma and D. patulo. Phytochcmistry 12: 683-687.
Tryon, A. and R. Tryon. 1974. Geographic patterns in temperate American ferns and some rela-
tionships in Thelypteris. Amer. Fern. 64: 99-104.
J.
Tryon, R. 1975. The Benjamin D. Gilbert fern herbarium. Amer. Fern 65: 60.
J.
Tryon, R. M. and G. Gastony. 1975. The biogeography of endemism in the Cyatheaceae. Fern
}.
Gaz. 11: 73-79.
Tryon, R. M. and D. S. Conant. 1975. The ferns of Brazilian Amazonia. Acta Amazon. 23-34.
5:
A
Tryon, R. 1976. revision of the genus Cyathea. Contr. Gray Herb. 206: 19-98.
Gastony, G. and R. M. Tryon. 1976. Spore morphology in the Cyatheaceae. The genera
II.
J.
Lophosoria, Metaxya, Spbaeropteris, Alsophila, and Nephelea. Amer. 738-758.
Bot. 63:
J.
Tryon, R. M. 1977, Studies on the American Cyatheaceae and on the classification of the family.
2839-2842.
Males. Bull. 30:
Fl.
Tkyon, M. and Vitale. 1977. Evidence antheridogen production and mediation
R. G. for of a
its
mating system in natural populations of fern gametophytes. Bot. Linn. Soc. 74: 243-249.
J.
Tryon, R. 1979. Biogeography of the Antillean fern flora. Pp. 55-68 in D. Bramwell (ed.) Plants
New
and Academic
islands. Press, York.
Tryon, A. R. Tryon, and Badre. 1980. Classification, spores, and nomenclature of the marsh
F., F.
Rhodora 461-474.
fern. 82:
Tryon, R. 1980. Review of Steyermark, A. and O. Huber, Flora of Avilo. Amer. Fern 70: 79.
J. J.
Tryon, R. M,, 1980. The ferns and fern allies of Minnesota, Ed. 2. Univ. Minnesota Press,
Jr.
Minneapolis.
Tryon, R. and A. Tkyon. 1981. Taxonomic and nomenclatural notes on ferns. Rhodora 83: 133-
137.
Tryon, R. 1982. Review of Smith, A. R., Pteridopbytes in D, E. Breedlove (ed.) Flora of Chiapas
Madrono
Port 2. 29: 216.
Tryon, R. and A. Tryon. 1982. Additional taxonomic and nomenclatural notes on ferns. Rho-
F.
dora 125-130.
84:
Tryon, R. M. and A. Tryon. 1982. Ferns and allied plants with special reference to tropical
F.
New
America. Springer-Verlag, York.
An new
Tryon, R. 1984. unusual Elaphoglossum from Peru. Amer. Fern 74: 108-110.
J.
Tryon, R. 1985. Fern speciation and biogeography. Proc. Roy. Soc. Edinburgh, B. 86: 353-360.
Tryon, R. 1986. Dicksoniaceae. Lophosoriaceae. Metaxyaceae. Cyatheaceae. Pp. 1-59 in G. Harling
and L. Andersson (eds.) Flora of Ecuador, No. 27. Swedish Research Councils, Stockholm.
Tryon, R. 1986. Some new names and combinations in Pteridaceae. Amer. Fern 76: 184-186.
J.
ROLLA MILTON TRYON,
9
JR.
Tryon, R. 1986. The biogeography of species, with special reference to ferns. But. Rev. [Lancaster]
117-156.
52:
Tryon, R. 1989. Ptcridophytes. Pp. 327-338 in H. Lieth and M. A. Werger (eds.) Tropical rain
J.
Amsterdam.
ecosystems.
forest Elsevier,
Ophioglossaceae-12. Cy-
Tryon, R. M. and R. G. Stolze. 1989. Pteridophyta of Peru. Part 1.
I.
New
1-145.
atheaceae. Fieldiana, Bot. Series 20:
Dennstaed-
Tryon, R. M. and R. G. Stolze. 1989. Pteridophyta of Peru. Part IT. 13. Pteridaceae-15.
New
1-128.
tiaceae. Fieldiana, Bot. Series 22:
Tryon, M., A. Tryon, and K. U. Kramer. 1990. Pteridaceae. Pp: 230-256, in K. Kubitzki (ed.)
R.
F.
and gymnosperms.
The families and genera of vascular plants. Vol. 1. Pteridopbyics Vol.
New
Kramer and Green. Springer-Verlag, York.
eds. K. U. S.
P.
Dryopteridaceae [with
Tryon, M. and Stolze. 1991. Pteridophyta of Peru. Part IV. 17.
R. R. G.
New
1-176.
collaboration of Mickel and R. C. Moran). Fieldiana, Bot. Series 27:
T.
J.
Thelypteridaceae (con-
Tryon, M. and Stolze. 1992. Pteridophyta of Peru. Part 16.
R. R. G. III.
New
1-80.
tributed by A, R. Smith]. Fieldiana, Bot. Series 29:
Aspleniaceae-21. Poly-
Tryon, M. and Stolze. 1993. Pteridophyta of Peru. Part V. 18.
R. R. G.
New
1-190.
podiaceae (with collaboration of B. Leon). Fieldiana, Bot. Series 32:
Tryon, M. and Stolze. 1994. Pteridophyta of Peru. Part VI. 22. Marsiliaceae-28. Isoctaccae
R. R. G.
New
1-123.
(with collaboration of R. Hickey and B. 01Igaard). Fieldiana, Bot. Series 34:
J.
Rhodora 335-343.
Tryon, 1997. Systematic notes on Oleandra. 99:
R.
name ebeneum Taxon 339-
Acrostichurn (Pteridaceae). 46:
Tryon, R. 1997. Proposal to reject the
340.
Churchill, H., R. Tryon, and D. S. Barrington. 1998. Development of the sorus in tree ferns:
1245-1252.
Dicksoniaceae. Canad. Bot. 76:
J.
North Amer-
Tryon, M. and A. Tryon. 1999. Observations on the phytogeography of eastern
R. F.
Pp 250-273 X-G Zang and K-H Shing Clung Memorial Volume. Institute
(eds.).
ican ferns. in
Chinese Acad. Beijing.
of Botany, Sci.,
428-438.
Tryon, 2000. Systematic notes on the Old World fern genus Oleandra. Rliodora 102:
R.
manual
and Moran, Fern grower's revised
Tryon, M. 2001. Review of Hoshizaki, B. R. C.
R.
J.
Rhodora
WiNDISCH
Bradea 267-276.
probable migration route and its present fern flora. 8:
